Plants moderately large, densely caespitose, golden yellow to golden brown, somewhat glossy. Stems creeping, densely or rarely loosely pinnate; hyalodermis absent, central strand present, weak; pseudoparaphyllia foliose, lanceolate, never filamentous. Stem and branch leaves with dorsal, lateral, and ventral stem leaves often different in shape and size. Stem leaves closely imbricate, narrowly lanceolate, plicate; base not or short-to long-decurrent; margins recurved at base, entire to subentire proximally, serrate, serrulate, or rarely entire distally; apex narrowly acute [blunt]; costa double; alar cells differentiated, subquadrate to rectangular; laminal cells smooth, . Branch leaves much smaller, narrower. Specialized asexual reproduction absent. Sexual condition dioicous; inner perichaetial leaves oblong-lanceolate, apex filiform-acuminate. [Seta reddish brown. Capsule inclined to horizontal, ovoid to cylindric, straight or weakly curved, not or weakly contracted below mouth; annulus little differentiated; operculum conic, sometimes short-rostrate; peristome double; exostome teeth with external surface transversely smooth to striolate proximally, papillose distally; endostome basal membrane 1/2 exostome teeth length, segments keeled, cilia 2-4. Calyptra naked. Spores rounded, finely papillose].