Plants small, in tomentose mats. Stems creeping, branches erect. Leaves contorted and spirally twisted around stem, ± undulate when dry, erect to wide-spreading when moist, lanceolate to lingulate, sometimes rugose; margins entire or denticulate; apex rounded-obtuse and mucronate or acute and subulate; costa percurrent or excurrent; basal laminal cells oblate to rounded; distal cells rounded-hexagonal or irregularly rounded to elliptic-hexagonal, 3-9 µm, smooth; marginal cells distinct from basal. Sexual condition dioicous or pseudautoicous; perichaetial leaves similar to stem leaves. Seta to 11 mm. Capsule exserted, oblong-ovate to cylindric, usually smooth or obscurely plicate when old, not constricted below mouth; stomata superficial; peristome rudimentary; exostome absent; endostome basal membrane usually low, papillose. Calyptra mitrate, conic, ± lacerate, or divided into many lobes, smooth, naked, plicate, covering less than 1/2 capsule. Spores isosporous or anisosporous.
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