Gymnocarpium dryopteris (L.) Newman

Oak fern (en), Gymnocarpium dryoptéride (fr), Polypode du chêne (fr), Lastrée du chêne (fr), Dryoptéride de Linné (fr), Gymnocarpium du chêne (fr), Dryoptéris de Linné (fr)

Species

Pteridophytes > Polypodiales > Cystopteridaceae > Gymnocarpium

Characteristics

Stems 0.5--1.5 mm diam.; scales 1--4 mm. Fertile leaves usually 12--42 cm. Petiole 9--28 cm, with sparse glandular hairs distally; scales 2--6 mm. Blade broadly deltate, 2-pinnate-pinnatifid, 3--14 cm, lax and delicate, abaxial surface and rachis glabrous or with sparse glandular hairs, adaxial surface glabrous. Pinna apex entire, rounded. Proximal pinnae 2--12 cm, ± perpendicular to rachis, with basiscopic pinnules ± perpendicular to costa; basal basiscopic pinnule usually sessile, pinnatifid or rarely pinnate-pinnatifid, if sessile then with basal basiscopic pinnulet often equaling or longer than adjacent pinnulet; 2d basal basiscopic pinnule sessile, with basal basiscopic pinnulet equaling or longer than adjacent pinnulet; basal acroscopic pinnule sessile, with basal basiscopic pinnulet longer than or equaling adjacent pinnulet. Pinnae of 2d pair usually sessile with basal basiscopic pinnule longer than or equaling adjacent pinnule and about equal to basal acroscopic pinnule; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule, often with entire, rounded apex. Pinnae of 3d pair sessile with basal basiscopic pinnule equaling adjacent pinnule and equaling basal acroscopic pinnules; basal acroscopic pinnule equaling or slightly shorter than adjacent pinnule. Ultimate segments of proximal pinnae oblong, entire to crenate, apex entire, rounded. Spores 34--39 µm. 2 n = 160.
More
Rhizomes slender, creeping, black, shiny, apex clothed with brown ovate-lanceolate scales. Fronds distant; fertile lamina (15-)20-30(-50) cm; stipe stramineous, 10-22(-35) cm, slender, with sparse scales at base; lamina nearly tripartite, usually 2-pinnate-pinnatifid, pentagonal-ovate or broadly ovate-triangular, 7-15(-20) cm long and wide, thinly herbaceous or submembranous, base broadly cuneate, apex acuminate; basal pair of pinnae nearly as large as other portion of lamina, narrowly triangular, (3.5-)5-9(-12) × 2.5-4(-7) cm, pinnate-pinnatifid, base subtruncate, with stalk (0.8-)1-1.5(-2.5) cm, portion of rachis between basal pinnae and central lamina ca. 3 cm; pinnules 5 or 6 pairs, oblong-lanceolate, 1.5-2(-4) × 0.5-2 cm, base rounded-cuneate, sessile, apex acute or acuminate, opposite or subopposite, spreading; largest pinnules with 6-10 pairs of segments, segments approximate, oblong to narrowly ovate, ca. 4 mm, lobed to narrow costular wing, entire to shallowly lobed at margin, rounded-obtuse at apex; second basal pair of pinnae 1.5-4 cm apart from basal pair, sometimes shortly stalked, upper pinnae sessile; veins pinnate in segment, simple, oblique, visible abaxially; rachis and costae slender, eglandular. Sori small, exindusiate, orbicular, abaxial on veins. Spore wall surface rugate, foveolate. 2n = 160.
Petiole 10–30 cm; blade yellow-green, scaleless, deltoid-pentagonal, to 18 × 25 cm, bipinnate-pinnatifid to tripinnate-pinnatifid, glabrous or occasionally slightly glandular especially along the rachis, pinnae several pairs, opposite, the members of the lowest pair deltoid and each nearly as long as the rest of the blade, with a petiolule to 2.5 cm long, distinctly asymmetrical, the lowest basiscopic segment evidently the largest, mostly a third as long as the main rachis or longer; ultimate segments oblong, 8–18 mm, obtuse, the margins crenate to subpinnatifid, not recurved; veins 2–6 pairs, mostly simple; 2n=160 (ours). Cool woods and talus-slopes; circumboreal, s. to Va., N.C., O., Ill., Io., and Ariz. A vegetatively reproducing hybrid with the next is G. ×heterosporum W. H. Wagner.
A fern.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality monoecy
Pollination hydrogamy
Spread anemochory
Mature width (meter) 0.5 - 1.0
Mature height (meter) 0.1 - 0.5
Root system creeping-root rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a temperate plant. In north China it grows between 300-2,900 m above sea level in damp areas in pine forests. It suits hardiness zones 4-8.
Light 1-4
Soil humidity 4-9
Soil texture 3-6
Soil acidity 2-6
Soil nutriment -
Hardiness (USDA) 6-9

Usage

Uses environmental use medicinal wood
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Gymnocarpium dryopteris habit picture by Terje Norli (cc-by-sa)
Gymnocarpium dryopteris habit picture by Yoan MARTIN (cc-by-sa)
Gymnocarpium dryopteris habit picture by Martin Bishop (cc-by-sa)

Leaf

Gymnocarpium dryopteris leaf picture by Robert Klink (cc-by-sa)
Gymnocarpium dryopteris leaf picture by Jukka Sjöblom (cc-by-sa)
Gymnocarpium dryopteris leaf picture by Terje Norli (cc-by-sa)

Distribution

Gymnocarpium dryopteris world distribution map, present in Albania, Andorra, Austria, Belgium, Bulgaria, Bosnia and Herzegovina, Belarus, Canada, Switzerland, China, Czech Republic, Germany, Denmark, Spain, Estonia, Finland, France, United Kingdom of Great Britain and Northern Ireland, Georgia, Greece, Greenland, Croatia, Hungary, Iceland, Italy, Japan, Kazakhstan, Liberia, Liechtenstein, Lithuania, Luxembourg, Latvia, North Macedonia, Malta, Montenegro, Mongolia, Netherlands, Norway, Poland, Korea (Democratic People's Republic of), Romania, Russian Federation, Saint Pierre and Miquelon, Serbia, Slovakia, Sweden, Turkey, Ukraine, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:17112110-1
WFO ID wfo-0001107317
COL ID 3HQVF
BDTFX ID 30719
INPN ID 100636
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Polypodium conjunctum Filix pumila Polypodium dryopteris Polystichum dryopteris Lastrea dryopteris Dryopteris dryopteris Currania dryopteris Dryopteris pumila Dryopteris pulchella Nephrodium dryopteris Aspidium dryopteris Carpogymnia dryopteris Phegopteris dryopteris Dryopteris linnaeana Thelypteris dryopteris Polypodium pulchellum Dryopteris triangularis Gymnocarpium dryopteris subsp. dryopteris Gymnocarpium dryopteris var. dryopteris Polypodium dryopteris var. glandulosum Polypodium dryopteris var. genuinum Polypodium dryopteris var. rigidius Polypodium dryopteris var. glabrum Gymnocarpium dryopteris