Gymnocarpium robertianum (Hoffm.) Newman

Scented oakfern (en), Lastrée du calcaire (fr), Polypode du calcaire (fr), Gymnocarpium de Robert (fr), Gymnocarpium du calcaire (fr)

Species

Pteridophytes > Polypodiales > Cystopteridaceae > Gymnocarpium

Characteristics

Stems 1--2 mm diam.; scales 2--4 mm. Fertile leaves usually 10--52 cm. Petiole 5--33 cm, with numerous glandular hairs distally; scales 2--6 mm. Blade broadly deltate, 2--3-pinnate-pinnatifid, 5--19 cm, usually firm and robust, abaxial surface moderately to densely glandular, rachis densely glandular, adaxial surface moderately glandular. Pinna apex acute. Proximal pinnae 3--13 cm, ± perpendicular to rachis, basiscopic pinnules ± perpendicular to costa; basal basiscopic pinnules either sessile or stalked, pinnate-pinnatifid or pinnatifid, if sessile then with basal basiscopic pinnulet usually shorter than adjacent pinnulet; 2d basal basiscopic pinnule sometimes stalked, if sessile then with basal basiscopic pinnulet shorter than or equaling adjacent pinnulet; basal acroscopic pinnule sometimes stalked, if sessile then with basal basiscopic pinnulet shorter than or equaling adjacent pinnulet. Pinnae of 2d pair usually stalked, if sessile then with basal basiscopic pinnule usually shorter than adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule shorter than adjacent pinnule, apex often entire, rounded. Pinnae of 3d pair usually sessile with basal basiscopic pinnule shorter than adjacent pinnule and equaling basal acroscopic pinnule; basal acroscopic pinnule equaling or shorter than adjacent pinnule. Ultimate segments of proximal pinnae oblong, entire to slightly crenate, apex entire, rounded. Spores 34--39 µm. 2 n = 160.
More
Much like no. 1 [Gymnocarpium dryopteris (L.) Newman] ; blade green, densely and finely glandular (as also the petiole), a little narrower, deltoid, to 14 × 16 cm; segments of the basal pinnae ovate-oblong, obtuse, sessile, the lowest basal one mostly less than a fourth as long as the main rachis; ultimate segments to 9 mm, with entire to crenate, mostly recurved margins; 2n=160 (ours). On calcareous rocks; circumboreal, s. to Pa., Mich., Io., and Ida. (G. jessoense; Dryopteris r.; Phegopteris r.)
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality monoecy
Pollination hydrogamy
Spread anemochory
Mature width (meter) -
Mature height (meter) 0.18 - 0.35
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

Light 1-5
Soil humidity -
Soil texture 1-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) 4-9

Usage

Uses medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Gymnocarpium robertianum habit picture by Justus Müller-Kiefer (cc-by-sa)
Gymnocarpium robertianum habit picture by Daniel Barthelemy (cc-by-nc)
Gymnocarpium robertianum habit picture by Daniel Barthelemy (cc-by-nc)

Leaf

Gymnocarpium robertianum leaf picture by Georgius (cc-by-sa)
Gymnocarpium robertianum leaf picture by Georgius (cc-by-sa)
Gymnocarpium robertianum leaf picture by Alain Lagrave (cc-by-sa)

Fruit

Gymnocarpium robertianum fruit picture by David Hocken (cc-by-sa)

Distribution

Gymnocarpium robertianum world distribution map, present in Albania, Andorra, Austria, Belgium, Bulgaria, Bosnia and Herzegovina, Belarus, Canada, Switzerland, China, Czech Republic, Germany, Algeria, Spain, Estonia, Finland, France, United Kingdom of Great Britain and Northern Ireland, Georgia, Greece, Croatia, Hungary, Italy, Japan, Kazakhstan, Liberia, Liechtenstein, Luxembourg, Latvia, North Macedonia, Montenegro, Netherlands, Norway, Poland, Romania, Russian Federation, Serbia, Slovakia, Sweden, Ukraine, United States of America, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:17112130-1
WFO ID wfo-0001109141
COL ID 3HQVR
BDTFX ID 30723
INPN ID 100640
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lastrea calcarea Gymnocarpium obtusifolium Phegopteris robertiana Polypodium obtusifolium Polypodium robertianum Lastrea robertiana Nephrodium robertianum Phegopteris robertianum Carpogymnia robertiana Currania robertiana Dryopteris robertiana Thelypteris robertiana Polypodium calcareum Aspidium robertianum Aspidium calcareum Phegopteris calcarea Gymnocarpium altaycum Lastrea obtusifolia Polypodium dryopteris var. robertianum Polypodium dryopteris var. calcareum Phegopteris dryopteris var. robertiana Gymnocarpium robertianum