Trees or tall shrubs, dioecious or monoecious. Young persistent branchlets similar to deciduous branchlets; all articles quadrangular; furrows shallow and open, exposing stomates. Leaves in whorls of 4. Male inflorescences on branchlets similar to vegetative branchlets, simple or appearing compound owing to condensation of branching; bracteoles persistent. Female inflorescences on short or elongated branchlets similar to vegetative branchlets. 'Cones' mostly borne amongst assimilatory branchlets; bracts much expanded laterally, broader than high; bracteoles protruding, rounded on back but neither split nor with a dorsal protuberance. Samara body striate, glabrous, yellow-brown or greyish, dull.
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