Plants dark green at least distally, usually forming a dense turf, occasionally loosely cespitose. Cauline leaves short-elliptic or ovate, occasionally long-elliptic, straight to weakly reflexed apically, 0.3-0.4(-0.6) mm, apex rounded to occasionally broadly acute, rarely apiculate, distal margins 1-stratose, distal laminal cells 9-10 µm in width, 1:1; distal adaxial costal cells elongate, occasionally short-rectangular. Specialized asexual reproduction usually present, of obovoid gemmae of 4-6 cells in leaf axils. Perichaetia single on an elongate axis, terminating the axis, or lateral on a short branch; perichaetial leaves stiff or flexuose, long-ligulate to ovate-lanceolate, basal cells rectangular to rhomboid, smooth, distal laminal cells quadrate and strongly papillose or similar to the basal cells. Capsule short-cylindric, wide-mouthed, exothecial cells bulging, usually 30-40 µm wide.