Plants scattered or very loosely cespitose. Stem to 0.4 cm, central strand absent. Leaves 0.25-1(-1.4) mm, erect to incurved when dry, spreading when moist, base broadened in lower 1/3-1/2, long-rectangular; margins entire; costa ending 2-4 cells below the apex, cells of adaxial surface elongate and flat (not bulging), costa in section with 1 or 2 stereid bands or costa homogeneous; distal laminal cells quadrate, with 3-4 irregular to 2-fid papillae, walls thin and evenly thickened, lumens angular; basal cells differentiated across leaf, filling the leaf base; medial cells hyaline to yellowish, smooth or weakly papillose above, 10-12 µm wide, 2-4:1, walls thin. Specialized asexual reproduction commonly present in sterile material as gemmae in the axils of leaves, elongate, irregularly clavate, short-branching, or spindle-shaped, of mostly 4-10 cells. Sexual condition dioicous; perichaetial leaves strongly differentiated and sheathing the seta. Capsule lacking peristome, annulus revoluble, operculum short-conic.
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