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Stamens 4, staminode absent, didynamous, subequal, usually exserted, filaments filiform, inserted subcentrally on corolla tube; anther cells diverging, at length divaricate.
Leaves opposite, petiolate, ovate or elliptic, dentate or subentire.
Calyx 3–5-lobed, subrotate or cup-shaped, persistent.
Style filiform, persistent, usually exserted.
Flowers subfascicled in axillary clusters.
Ovary bilocular, ovoid; ovules numerous.
Seeds compressed, narrowly winged.
Shrubs or small trees, glabrous.
Pedicels bibracteolate.
Fruit a berry.
