Shrubs or trees, monoecious or dioecious; indumentum of simple hairs and gland-tipped hairs or sessile peltate-stellate hairs with central cells. Leaves [alternate or] opposite; stipules present, sometimes interpetiolar; leaf blade simple, base not glandular, margin entire, pinnately or palmately 3-veined. Inflorescences terminal or axillary, sometimes ramiflorous, [branched or] unbranched, male flowers 1-3 per bract, female 1 per bract. Male flowers: sepals 2-4, valvate; petals absent; disk absent; anthers 2-locular, extrorse, longitudinally dehiscent; pistillode absent. Female flower: sepals (3 or)4-6(or 7), imbricate or valvate; petals absent; disk absent; ovary [3-locular] with 1 ovule per locule; styles short; stigmas erect, adaxially shortly papillose. Fruit a capsule, spiny, spines sometimes gland-tipped. Seeds ± globose, 3-angled, without aril or carunculate.
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