Heliotropium ciliatum Kaplan

Species

Angiosperms > Boraginales > Boraginaceae > Heliotropium

Characteristics

Erect rhizomatous perennial herb up to 80 cm high. Stem usually branched from the base, somewhat swollen below the branches insertions, angular as for the branches with short appressed hairs. Leaves usually sparse; petiole 2-20(25) mm long, with hair-covering like the branches; lamina 20-50 x 5-20 mm, lanceolate or ovate, with hard short appressed hairs above and on the nerves below, covered with the swollen bases of minute hairs below, rarely roughish on both surfaces, acute to obtuse and sometimes mucronate at apex, acute to obtuse, decurrent on the petiole and often asymmetrical at base, margins usually repand and undulate. Cymes lax, up to 19 cm long when completely expanded, usually 2, rarely single, terminal; common peduncle 1-4 cm long. Flowers sessile or subsessile. Calyx minute, callose-strigose or subglabrous outside, subglabrous or glabrous inside; lobes 0.5-1.0(2.0) mm long, ovate to ovate-triangular, acute, callose-ciliate mainly at the base, thick. Corolla hypocrateriform, white, sparsely strigose outside on the upper part, otherwise glabrous outside and inside; tube 3.5-5.0 mm long, slightly widened in the upper half; limb 3-5(6) mm in diam.; lobes 1.2-2.2(2.5) mm long, triangular-acuminate, usually with undulate and ± crenate margins, bent inwards in bud, spreading. Stamens inserted about the middle of the corolla tube; anthers 1.2-1.5 mm long, narrowly oblong, emarginate at apex, obtuse to subcordate at base, subsessile. Ovary glabrous. Style 1.0-1.5(2.0) mm long, filiform, glabrous; stigmatic ring not very well marked; sterile appendix 1.0-1.4 mm long, narrowly conical, deeply 2-lobed, hirtellous. Fruits c. 4 x 5 x 4 mm, saddle-shaped at apex, tuberculate-crested, glabrous; nutlets connate in pairs, each pair 2-horned, 2-seeded, with the inner surface flat.
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Perennial herb, much branched from a woody base, 0.2-0.8 m high. Stems erect. Leaves petiolate; blade ovate to elliptic, 10-30 x 2-10 mm, slightly decurrent, base cuneate, apex acute, margin undulate, shallowly toothed, lower leaf blade with bulbous-based setae, upper part of seta short and stout; petioles 3-12 mm long. Flowers: in helicoid cymes, ebracteate; upper part of style head cylindrical; calyx pubescent, persistent, deeply lobed, much shorter than corolla, lobes triangular; corolla white, throat orange, tube cylindrical, ± 3.5 mm long, appressed pubescent in upper part, lobes acuminate, ± 1.5 mm long; Oct.-May. Fruit 2 nutlets, rugose, tuberculate-crested, saddle-shaped at apex, connate in pairs.
Leaves usually sparse; petiole 2–20(25) mm. long, with hair-covering like the branches; lamina 20–50 x 5–20 mm., lanceolate or ovate, with hard short appressed hairs above and on the nerves below, covered with the swollen bases of minute hairs below, rarely roughish on both surfaces, acute to obtuse and sometimes mucronate at apex, acute to obtuse, decurrent on the petiole and often asymmetrical at base, margins usually repand and undulate.
Corolla hypocrateriform, white, sparsely strigose outside on the upper part, otherwise glabrous outside and inside; tube 3.5–5.0 mm. long, slightly widened in the upper half; limb 3–5(6) mm. in diam.; lobes 1.2–2.2(2.5) mm. long, triangular-acuminate, usually with undulate and ± crenate margins, bent inwards in bud, spreading.
Perennial herb, much branched from a woody base, 0.2-0.8 m high. Stems erect. Leaves petiolate; blade narrowly ovate to ovate; petioles 3-12 mm long. Flowers in helicoid cymes. Calyx much shorter than corolla. Corolla white, throat orange. Flowering time Oct.-May. Fruit of glabrous, rugose nutlets.
Erect perennial herb, up to 0.8 m high. Hairs on stem appressed. Calyx lobes thick, callose. Corolla lobes acuminate, tube long. Fruit tuberculate-crested, saddle-shaped at apex; nutlets connate in pairs. Flowers white.
Calyx minute, callose-strigose or subglabrous outside, subglabrous or glabrous inside; lobes 0.5–1.0(2.0) mm. long, ovate to ovate-triangular, acute, callose-ciliate mainly at the base, thick.
Style 1.0–1.5(2.0) mm. long, filiform, glabrous; stigmatic ring not very well marked; sterile appendix 1.0–1.4 mm. long, narrowly conical, deeply 2-lobed, hirtellous.
Fruits c. 4 x 5 x 4 mm., saddle-shaped at apex, tuberculate-crested, glabrous; nutlets connate in pairs, each pair 2-horned, 2-seeded, with the inner surface flat.
Stamens inserted about the middle of the corolla tube; anthers 1.2–1.5 mm. long, narrowly oblong, emarginate at apex, obtuse to subcordate at base, subsessile.
Stem usually branched from the base, somewhat swollen below the branches insertions, angular as for the branches with short appressed hairs.
Cymes lax, up to 19 cm. long when completely expanded, usually 2, rarely single, terminal; common peduncle 1–4 cm. long.
Erect rhizomatous perennial herb up to 80 cm. high.
Flowers sessile or subsessile.
Ovary glabrous.
Life form perennial
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.5 - 0.8
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

In riverbanks, savannahs with trees and shrubs and in dry woodlands, on sandy or rocky soils; at elevations up to 1,050 metres.
Light -
Soil humidity 1-3
Soil texture 6-8
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses medicinal
Edible -
Therapeutic use Headache (unspecified), Antidote(Reptile) (unspecified), Lactagogue (unspecified), Eye (unspecified), Gonorrhea (unspecified), Bite(Scorpion) (unspecified), Bites and stings (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) 14 - 42
Germination temperacture (C°) 18 - 23
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Heliotropium ciliatum unspecified picture

Distribution

Heliotropium ciliatum world distribution map, present in Botswana, Mozambique, Namibia, eSwatini, South Africa, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:116749-1
WFO ID wfo-0001215449
COL ID 3KCCT
BDTFX ID -
INPN ID -
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Heliotropium tuberculosum Heliotropium gibbosum Heliotropium pustulatum Tournefortia tuberculosa Heliotropium ciliatum