Plants small, caespitose, greyish green to blackish green dry, yellowish green wet, reddish brown below; terricolous. Stems 2-4 mm tall, unbranched; in section with large central strand, cortical cells lax, single outer row of cells smaller, incrassate, red-brown. Leaves appressed, spirally twisted dry, patent wet; oval to lingulate, 1.0-2.6 mm long; apex obtuse; margins plane, entire, marginal cells differentiated, forming unistratose border 3-5 cells wide. Costa mucronate; ventral superficial cells quadrate, papillose, dorsal superficial cells rectangular, thickened, with scattered, low, blunt papillae; in section with 2-4 guide cells, ventral cells in 1-2 rows, large, surface cells papillose, dorsal stereid band 4-5 cells thick, dorsal surface cells substereids, 1-2 lateral surface cells similar to ventral surface cells. Upper laminal cells quadrate to hexagonal, some transversely rectangular, with 4-5 low, C-shaped papillae over lumen; marginal cells quadrate to rectangular, some transversely rectangular, incrassate, smooth to weakly papillose, generally forming distinct border above base; basal cells quadrate to rectangular, smooth, smaller toward margin. Dioicous. Perichaetia terminal, leaves undifferentiated. Seta 10-12 mm long, yellow to red; capsule erect, cylindrical, 2.0-2.5 mm long, red-brown; peristome teeth divided to base into 2 long papillose filaments, 0.8 mm long, spirally twisted one turn; operculum conical, 1.0 mm long; spores round, to reniform, 10-12 µm, smooth.
Home