Tree 6-30 m. Twigs 2.5-5 mm diameter, early glabrescent, hairs greyish brown, 0.1 mm long or less; bark finely to coarsely striate, not flaking; lenticels usually conspicuous. Leaves membranous or chartaceous, (elliptic-)oblong, 15-27 by 5-10 cm, base attenuate, apex acute(-acuminate), sometimes ± blunt; upper surface drying olivaceous to dark brown, lower surface glabrous; dots absent; midrib ± flat above; nerves 9-18(-20) pairs, thin, flat or slightly raised above, lines of interarching indistinct; venation lax, faint on both surfaces; petiole 5-15 by 2-3 mm; leaf bud 12-20 by 2-3 mm, with dense grey-brown to rusty hairs 0.1 mm long or less. Inflorescences behind the leaves, with sparse (subsp. kinabaluensis) or dense hairs 0.1 mm long; in male: rather stout, about 3 (or 4) times branched, many-flowered, 5-14 by 3-10 cm, peduncle (0.6-)l-3 cm long; female 2 (or 3) times branched, 2-7 by 1.5-4 cm, fewer flowered than male; bracts ellipsoid to oblong, acutish, densely short-pubescent, 2-5 mm long, caducous; flowers in male in loose clusters of 2-5, perianth 3-(or 4-)lobed, glabrous, pedicel glabrous (subsp. kinabaluensis) or partly thinly pubescent (subsp. subalpina), not articulated. Male flowers: pedicel 1.5-2 mm, slender; buds broad-ellipsoid or subglobose, 1.6-2.3 mm long, cleft c. 1/2, lobes 0.2(-0.3) mm thick; androecium globose or broadly ellipsoid, 1.5 mm long, cross section circular (Plate 3: 84); thecae 16-24, almost completely sessile, free apices 0.1 mm, curving towards the apex and concealing the narrow apical hollow, 0.3-0.5 mm deep, of the broad column; androphore narrow, 0.2-0.3 mm long, largely hidden by the anthers. Female flowers: pedicel 1.5 mm long; buds ellipsoid, 2.5 by 1.8-2.1 mm, glabrous (subsp. kinabaluensis), cleft (1/3—)l/2; ovary subglobose to broadly ellipsoid, 1.2-1.5 mm long, glabrous, stigma minutely 2-lobed, 0.1 mm high. Fruits 2-6 per infructescence, subglobose, broadly ellipsoid, or ellipsoid-oblong, 2.5-5 cm long, glabrous; fruiting pedicel 3-7 mm long; perianth not persistent.
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