Liana to 40 m, rarely small tree to 3 m, or climbing shrub, up to 10 cm diam. Climbing shoots with first internodium (2.5—)6—18 cm, following internodia very short with some distichously arranged, reduced, linear to orbicular leaves, a few of these shoots with a long-peduncled, reduced inflorescence or a climbing hook, or the climbing shoot above the hooks growing out to a leafy and often fertile branch. Hooks distichously or subopposite, woody, (3.5-) 6.5-8.5 cm long, apically often with bract-like organs, peduncles, cymes, or reduced leaves. Stipules entire, obtuse to acute, with or without glandular teeth, persistent or sometimes caducous, sometimes only present as a glandular tooth, subulate or triangular, 0.5-0.8 by 0.2-0.8 mm. Leaves glabrous (rarely some hairs on midrib), narrowly elliptic to obovate, sometimes broadly elliptic or ovate, sometimes asymmetric, (6-)ll-27(-36) by (1.6-)3-9.5 (-11) cm; base cuneate, seldom rounded; margin often wavy, glandular-crenulate to crenate, apex acute to acuminate, rarely obtuse, acumen up to 2 cm; nerves 7-16 pairs, slightly arching upwards and almost reaching the margin, meeting in an often somewhat obscure intramarginal vein or in the dense reticulatron; petiole 3-20 mm. Inflorescence an axillary or terminal compound panicle, sometimes a raceme; terminal panicles often leafy, cone-shaped to obovoid, (4-)12-18(-32) by (2.5-)1.5-14 cm with 5-25(-30) more or less closely set subinflorescences, sometimes up to 5(-8) pseudoterminal panicles together; axillary panicles patent, narrowly obovoid to broadly ellipsoid, (2—)6—11(—12) by (l-)2-6.5 cm, with (4-) 10-23 subinflorescences; peduncle up to 6.5 cm; cymes up to 15, 0-5 times branched; bracts persistent, obtuse deltoid, (1.5-)2.5-5 by 0.2-1 mm, with 2-6 glandular teeth. Pedicel articulate and often thickened below the flower, 1.5-3 mm. Bracteoles like the bracts but smaller. Flowers heterodis-tylous, 7-9 by 5-9 mm. Sepals ± orbicular to transversely broadly elliptic, (1.8-)2-3(-4) by 2-4(-4.5) mm, shortly connate to free, appressed but patent in fruit, outer 2 smaller and thicker. Petals recurving during anthesis, narrowly elliptic to-obovate, 6-9.5 by 1.5 —3(—4) mm; base often indistinctly 0.8 mm clawed. Stamens shortly persistent, obdiplostemon-ous, in short-styled flowers (4.5-)4.8-6 and (5—)5.5—7.5 mm, in long-styled flowers (2.5-) 3.8-4.5(-4.8) and (3.8-)4.2-5(-5.8) mm; tube 1 —1.5(—2) mm, partly persistent in fruit; base often irregularly thickened or marked by darker coloured semi-ellipsoid, sometimes apically notched, staminal glands at the base of long or short stamens, these later becoming connate with each other; filaments triangular at base, apically filiform. Ovary 5-loculed, ovoid to broadly ellipsoid, 1-2 by 0.8-1.2 mm; styles 5, basally shortly connate and apically (in short-and long-styled forms) hooked or sigmoidly curved or (in long-styled forms) slightly arching to curving downwards, sometimes of different length, in short-styled flowers (1—)1.5—2.8 mm, in long-styled flowers (3-)3.2-3.5(-5.2) mm; stigma 2-lobed. Drupe 15-ribbed, ovoid to subglobose, (8—)10—13 by (7—)8-10.5 mm, splitting into 5 pyrenes; mesocarp fleshy, surrounding the pyrenes, between the pyrenes often disappearing thus giving rise to a false locule; endocarp bony, thin, 3-ribbed, ribs branching, the branches connected by bony septa, forming cavities filled with aerenchyma; septa and aerenchyma dissolving when ripe. Seed 1 or 0 in each pyrene, ventrally-apically attached by a short funicle, medially flattened, asymmetrically semi-ellipsoid, c. 8.5 by 3.5 mm; hilum and arillus not obvious; testa thin, 2-layered. Embryo c. 8 by 3 mm; radicle flattened, arched, c. 1.5 by 2.5 mm. Endosperm fleshy.
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Scandent shrub or liane to 40 m climbing by woody subopposite hooks, sometimes a tree to 3 m. Leaves elliptic to obovate, 5–35 cm long, usually acute to acuminate, base cuneate to rounded, margin glandular-crenulate to crenate. Inflorescence a panicle or raceme 4–32 cm long. Flowers fragrant. Sepals orbicular to elliptic, 2–4 mm long, ciliolate. Petals obovate to elliptic, 6–10 mm long, recurved, yellow to orange. Stamens exserted; longer filaments to 8 mm long. Ovary 5-locular; styles 5. Fruit ovoid, 10–13 mm long, ribbed, yellow to brown. Seeds 8–9 mm long.