Humulus lupulus L.

Hop (en), Houblon (fr), Houblon grimpant (fr), Houblon lupulin (fr), Vigne du Nord (fr)

Species

Angiosperms > Rosales > Cannabaceae > Humulus

Characteristics

Perennial twining climber with stems to 6 (–8) m long and scabrid with reflexed hairs. Leaves opposite, with lamina ovate-cordate to ± circular, 5–13 (–25) cm long and about as wide, unlobed or more usually deeply 3–7-lobed, crenate-serrate; upper surface with appressed hairs with squamellate bases or only the bases remaining; lower surface paler, bearing yellowish, resinous, bitter and aromatic granules; petiole to 12 cm or more long. Peduncle slender, scabrous. Female inflorescence cone-like, 2–5 cm long at maturity; flowers subtended by large, persistent bracts; bracts ovate-acute, pale green to light brown, with aromatic granules, yellow-amber at maturity; outer surface of bracts pubescent or with appressed hairs outside, prominently veined inside. Seed c. 3 mm long, 2 mm wide, acute, with a central rib, shiny, brown, enclosed by a single large perianth bract.
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Herbs , perennial, rhizomatous, 1-6(-7) m. Stems branched. Leaves: petioles usually shorter than blades. Leaf blade ± cordate, palmately 3-7-lobed, sometimes unlobed, 3-15 cm, margins dentate-serrate; surfaces abaxially with veins glabrous or with soft pubescence, without straight, erect hairs, glands yellow, adaxially margins of younger leaf blades with few or no cystolithic hairs. Inflorescences: staminate with flowers whitish to yellowish, anthers glandular; pistillate usually racemes, 10-20 mm, pedunculate; bracteole margins not ciliate-hairy. Infructescences pendulous, pale yellow, conelike, ovoid to oblong, (1-)2-3(-6) cm; bracteoles with yellow glands. Achenes yellowish, ovoid, compressed, glandular. 2 n = 20, including 2 or more sex-determining chromosomes.
Dioecious or monoecious climber, with extensive roots. Stems striate, up to 10 m long and clothed in rough deflexed hairs. Lvs broadly ovate, not lobed or deeply 3-5-lobed; lobes acute to acuminate, serrate, ± glabrous to sparsely hairy above, ± glabrous to moderately hairy with yellow glandular hairs below. Petiole usually ± = lf blade. Stipules 2 per node and 2-fid or 4 per node and entire. Bracts of ♀ fls triangular, hairy, green, elongating and becoming yellowish at fruiting to form a cylindric-ovoid infructescence (the hop) with yellow glandular hairs especially at base of bracts. Calyx of ♂ fls ovate-triangular, green, hairy. Achene ovate-ellipsoid, covered by glandular calyx.
Plants perennial. Petiole usually shorter than leaf blade; leaf blade 3-5(-7)-lobed, sometimes simple, 4-11 × 4-8 cm, abaxially glabrous or with scattered soft pubescence but without rigid spinulose hairs on veins, adaxially with few or no cystolith hairs marginally when young, base cordate, margin coarsely serrate, apex acute. Female flowers 2 per bract at least in middle of inflorescence; bracts imbricated into a globose spike. Infructescences globose, 3-4 cm in diam.; bracts ovoid, 1.5-2 cm, dry, membranous, apex acute. Achenes flat, included in bracts. Fl. autumn.
Rhizomatous perennial vine to 10 m; principal lvs as broad as long, cordate at base, 3-lobed to below the middle, the upper lvs often broadly ovate and lobeless; petiole shorter than the blade; staminate infls 5–15 cm; pistillate spikes 1 cm, becoming 3–6 cm, the persistent, accrescent bracts entire and mostly blunt, stramineous, very glandular at base; 2n=20. Moist soil; N.S. to Man., Mont., and Calif., s. to N.C., Ark., and Ariz., and widely distributed in the Old World. July, Aug. 5 geographically significant vars., 3 in our range.
A climbing twiner which keeps growing from year to year. It can be 4.5-7.5 m long. The stems are rough. The leaves are deeply lobed and light green. They are 10-15 cm long. There are teeth around the edge. The stems are square, thin and bristly. The flowers are green and cone-like. There are several named varieties.
Life form perennial
Growth form herb
Growth support climber
Foliage retention -
Sexuality dioecy
Pollination anemogamy
Spread anemochory
Mature width (meter) -
Mature height (meter) 5.0 - 6.0
Root system creeping-root fibrous-root rhizome
Rooting depth (meter) 1.5
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a temperate plant. They prefer moist but well-drained soil. Plants are frost hardy. It cannot tolerate salty seaside conditions. They do best with a pH of 5.5-7. In the Himalayas it grows between 3,000-3,700 m above sea level. It suits hardiness zones 5-10. Hobart Botanical Gardens.
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Hedgerows, woodlands and sunny waste ground.
Light 4-9
Soil humidity 1-7
Soil texture 1-6
Soil acidity 4-7
Soil nutriment -
Hardiness (USDA) 5-8

Usage

The overlapping bracts which cover the female flower are used in brewing. They flavour beer and act as a preservative. Hops can also be used to make bread rise. They are dried and the oil has a smell. The flowers are used in bread. The young shoots are cooked and eaten. They are stewed, used in soups and also used in omelettes. Caution: Hops can cause mild depression.
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Has a long record of human usage, mostly in the brewing industry (the infructescences are used in brewing), but also in the past in bread-making and for its herbal properties. Many cultivars are known, the commonest in Australia being 'Pride of Ringwood' (Walsh 2015).
Uses beer breeding dye environmental use essential oil fiber fodder food gene source leaf vegetable material medicinal oil perfumery pharmaceutical wood
Edible flowers leaves rhizomes roots seeds shoots
Therapeutic use Analgesics (flower), Anesthesia (flower), Anthelmintics (flower), Anticonvulsants (flower), Anti-infective agents, local (flower), Antioxidants (flower), Appetite stimulants (flower), Astringents (flower), Diuretics (flower), Hypnotics and sedatives (flower), Hypothermia (flower), Inflammation (flower), Muscle relaxation (flower), Nervous system diseases (flower), Neuralgia (flower), Parasympatholytics (flower), General tonic for rejuvenation (flower), Sleep aids, pharmaceutical (flower), Sleep initiation and maintenance disorders (flower), Analgesic (fruit), Febrifuge (fruit), Gastrointestinal Aid (fruit), Anxiety disorders (fruit), Diarrhea (fruit), Diuretics (fruit), Anti-bacterial agents (leaf), Appetite stimulants (leaf), Dermatitis (leaf), Diabetes mellitus (reproductive organ), Sedative (root), Dermatological Aid (root), Analgesic (unspecified), Antirheumatic (Internal) (unspecified), Breast Treatment (unspecified), Gynecological Aid (unspecified), Kidney Aid (unspecified), Sedative (unspecified), Urinary Aid (unspecified), Ear Medicine (unspecified), Stimulant (unspecified), Toothache (unspecified), Tonic (unspecified), Diuretic (unspecified), Dermatological Aid (unspecified), Pulmonary Aid (unspecified), Anaphrodisiac (unspecified), Antiseptic (unspecified), Astringent (unspecified), Bitter-Principle (unspecified), Boil (unspecified), Cancer (unspecified), Dyspepsia (unspecified), Fever (unspecified), Hypnotic (unspecified), Nerves (unspecified), Nervine (unspecified), Poultice (unspecified), Rheumatism (unspecified), Shampoo (unspecified), Soporific (unspecified), Stomachic (unspecified), Sudorific (unspecified), Tea (unspecified), Tumor(Liver) (unspecified), Vermifuge (unspecified), Anodyne (unspecified), Cystitis (unspecified), Debility (unspecified), Inflammation (unspecified), Tuberculosis (unspecified), Analgesics (unspecified), Anthelmintics (unspecified), Anti-bacterial agents (unspecified), Antirheumatic agents (unspecified), Asthenia (unspecified), Cosmetics (unspecified), Dermatomycoses (unspecified), Digestive system diseases (unspecified), Diuretics (unspecified), Estrogen receptor modulators (unspecified), Hypnotics and sedatives (unspecified), Nervous system diseases (unspecified), Parasympatholytics (unspecified), General tonic for rejuvenation (unspecified), Sleep aids, pharmaceutical (unspecified), Diaphoretic (unspecified), Depurative (unspecified)
Human toxicity weak toxic (fruit)
Animal toxicity -

Cultivation

Plants are grown from semi-ripe cuttings. Plants are placed 2.5 m apart and need a structure to climb. Sometimes plants are grown from seeds if the cultivars set seed.
Mode cuttings divisions seedlings
Germination duration (days) 25 - 30
Germination temperacture (C°) 13 - 14
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -20
Optimum temperature (C°) 15 - 19
Size -
Vigor -
Productivity -

Images

Habit

Humulus lupulus habit picture by Christiane Mumme (cc-by-sa)
Humulus lupulus habit picture by Krzysztof Golucki (cc-by-sa)
Humulus lupulus habit picture by Émile Maurice (cc-by-sa)

Leaf

Humulus lupulus leaf picture by Oliver Engelen (cc-by-sa)
Humulus lupulus leaf picture by Christiane Mumme (cc-by-sa)
Humulus lupulus leaf picture by Simone Riva (cc-by-sa)

Flower

Humulus lupulus flower picture by Remi Espert (cc-by-sa)
Humulus lupulus flower picture by Simone Roch (cc-by-sa)
Humulus lupulus flower picture by Francois Lelievre (cc-by-sa)

Fruit

Humulus lupulus fruit picture by Helene Vialle (cc-by-sa)
Humulus lupulus fruit picture by Syd Paul Labartette (cc-by-sa)
Humulus lupulus fruit picture by Pascal Huguenin (cc-by-sa)

Distribution

Humulus lupulus world distribution map, present in Afghanistan, Albania, Argentina, Armenia, Australia, Austria, Azerbaijan, Belgium, Bulgaria, Bosnia and Herzegovina, Belarus, Brazil, Canada, Switzerland, China, Czech Republic, Germany, Denmark, Spain, Estonia, Finland, France, United Kingdom of Great Britain and Northern Ireland, Georgia, Greece, Croatia, Hungary, India, Iran (Islamic Republic of), Italy, Japan, Kazakhstan, Kyrgyzstan, Korea (Republic of), Liberia, Liechtenstein, Lithuania, Luxembourg, Latvia, Morocco, Moldova (Republic of), North Macedonia, Myanmar, Montenegro, Netherlands, Norway, New Zealand, Pakistan, Poland, Portugal, Romania, Serbia, Slovakia, Slovenia, Sweden, Syrian Arab Republic, Turkey, Ukraine, United States of America, Uzbekistan, Viet Nam, and South Africa

Identifiers

LSID urn:lsid:ipni.org:names:303502-2
WFO ID wfo-0000725866
COL ID 6MFNY
BDTFX ID 34958
INPN ID 103031
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lupulus communis Lupulus amarus Humulus volubilis Humulus vulgaris Lupulus humulus Lupulus scandens Humulus lupulus var. cordifolius Humulus lupulus var. lupulus Humulus lupulus