Hydrostachys polymorpha Klotzsch

Species

Angiosperms > Cornales > Hydrostachyaceae > Hydrostachys

Characteristics

Leaves narrowly elliptic, linear-oblong or lanceolate in outline, 15–75(–90) cm. long, 2–3-pinnate; petiole 5–25 cm. long, verrucose at base, the emergences becoming largest at base of limb.. Rhachis bearing broadly elliptic emergences 2 × 2 mm., with much smaller ones between them.. Primary pinnules 3–7 cm. long, 15–50 on each side of the primary rhachis, covered with elliptic emergences.. Secondary pinnules 0.4–2 cm. long, entirely covered with simple linear to elliptic emergences.. Inflorescences pinkish, grouped on the rhizome, often numerous (up to 50), stiffly erect.. ♀ 10–20 cm. long including the 5–10 cm. long peduncle, covered with small emergences, the spike itself 5–10 cm. long, with 150–200 flowers; bracts 2–3 mm. long, elliptic, entire at the apex, the spur with 2–3 apical rows of emergences, usually fused among themselves and giving the bract a wrinkled appearance; ovary flanked by 2 bundles of hairs; stigmas crimson.. ♂ inflorescence 10–30 cm. long including the verrucose 5–20 cm. long peduncle, the spike itself 5–10 cm. long with 100–200 flowers; bract irregularly shaped, 2 × 2 mm., the spur with 3–4 rows of emergences ± fused between themselves particularly the upper pair of rows. Stamens subsessile.. Fig. 1/1–8.
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Aquatic fern-like herbs up to 40 cm tall. Leaves submerged, basal, 10-40 cm long, 2-3 pinnatisect, covered all over with warty to ligulate excrescences; petiole and rhachis flattened, sometimes winged; pinnae sub-opposite, up to c. 3 cm long, numerous, spreading (somewhat resembling a fox-tail), pinnules much ramified, irregular, filamentous (the so-called 'gill-tufts'). Spikes emergent, erect, dioecious; male and female sub-similar, up to 30 cm long, arising from the base of the plant; scape verrucose or rarely smooth, stout; male flowers with the imbricate bracts broadly obovate, verruculose near the obtuse apex, margin recurved, locules joined only at the base; female flowers with conchiform bracts, c. 3 mm long, apex lip-like, central nerve decurrent below. Capsule hidden inside the bract, compressed-obovoid, smooth; seeds minute, orange.
Pinnae in c. 25–70 pairs, alternate to subopposite, 7.5–50 mm long, tapering-cylindric to ellipsoid in outline, those of at least some leaves differentiated into an upper and lower surface, with scale-like emergences all around the pinna axis; emergences simple, entire, ± closely spaced on the upper surface, and variously lobed, laciniate or ± filiform-branched on the lower surface, or all pinna emergences simple and scale-like; pinnules (pinna branches) sometimes present on the lower surface, 4–10 mm long, with linear to elliptic emergences.
Male inflorescences 3.5–22 cm long; peduncles 1.5–15 cm long, covered with emergences; bracts 1.5–2 ? 1.5–2.0 mm, irregularly shaped, with several rows of tubercles often ± fused into ridges, apical lip entire and rounded or sometimes recurved, the base decurrent. Anther thecae c. 0.5 mm long, elliptic to rounded, joined at base only, divergent.
Bracts of female inflorescence 2–4 ? 1.5–2 mm, conchiform, rounded to broadly elliptic in outline, smooth to verruculose, with 1–4 rows of tubercles towards the apex sometimes fused into ridges; the apical lip flange-like, rounded, entire, recurved; the central nerve decurrent below, lateral nerves sometimes prominent.
Leaves fern-like in appearance, few to numerous from the rootstock apex, trailing, submerged, up to 70 cm long, narrowly elliptic to lanceolate in outline, often 2–3-pinnate, usually with a differentiated upper and lower surface.
Petiole and rhachis ± flattened, bearing numerous small ovate to ligulate scale-like emergences; emergences all similar in shape but varying in size and largest on the rhachis, the largest up to 3 ? 2 mm; petiole 4–25 cm long.
Female inflorescences 6–30 cm long; peduncles 3–17 cm long, covered with tiny emergences, (sometimes flaking off on dried specimens to give smooth surface);spikes 150–200-flowered.
Inflorescences emergent, few–numerous (up to c. 50), scapose, spicate above, pendulous in bud, stiffly erect at anthesis.
Ovary 1.5–2 ? 1–1.5 mm, ellipsoid, glabrous, usually with a bundle of hairs on each side; styles 1–2.0 mm long, glabrous.
Aquatic caespitose perennial herb from a ± globose woody base, firmly attached to rocks by stiff radiating roots.
Fruit 2–2.5 ? 0.75–1 mm, elliptic, glabrous.
Seeds ovoid to oblong.
Life form perennial
Growth form herb
Growth support aquatic
Foliage retention -
Sexuality dioecy
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.4
Root system rhizome
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months -
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses medicinal
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Hydrostachys polymorpha world distribution map, present in Angola, Mozambique, Malawi, Namibia, Tanzania, United Republic of, South Africa, and Zimbabwe

Conservation status

Hydrostachys polymorpha threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:688228-1
WFO ID wfo-0000726742
COL ID 6ML7V
BDTFX ID -
INPN ID -
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Hydrostachys polymorpha Hydrostachys cristulata Hydrostachys multipinnata Hydrostachys natalensis