Plants small to large, yellow to dark green or blackish. Stems irregularly and often freely branched; hyalodermis absent, central strand usually present; paraphyllia often present, ; rhizoids on stem, never from abaxial costa surface, often forming tomentum, sparsely branched, smooth; axillary hairs , distal cells 1 or 2, hyaline. Stem and branch leaves erect, not or slightly falcate-secund, oblong-lanceolate to oblong-ovate, rarely broadly ovate, not plicate, to or longer than 1 mm; base not decurrent; margins plane, entire to denticulate, limbidia absent; apex gradually acuminate, acute, or obtuse, ; costa single, ending mid leaf to excurrent, broad, often curved beyond mid leaf; alar cells not or poorly differentiated, rectangular to quadrate, not or slightly inflated, green to brownish orange, , region indistinct, ovate, along basal margins; medial laminal cells rhombic to oblong-hexagonal, . Sexual condition autoicous. Capsule inclined, cylindric to oblong-cylindric, arcuate; peristome perfect; exostome teeth bordered; endostome cilia in groups of 1-3, nodulose to appendiculate. Spores 12-45 µm.
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