Plants soft, pale yellow, brownish yellow-green, or bright green. Stems to 5 cm, leafy throughout, irregularly branched; hyalodermis absent, epidermal cells small, walls thick, similar to subadjacent cortical cells, central strand hyaline to brownish. Leaves appressed-imbricate and plane or erect-spreading and convoluted-ruffled parallel to long axis of leaf, straight, sometimes secund, spreading when dry or moist, oval or orbicular, sometimes broadly ovate, concave, (1-)1.2-1.6(-1.8) × (0.9-)1-1.7(-1.8) mm; margins plane, entire, less often finely denticulate or serrulate in apex; apex broadly acute to short-apiculate; costa slender, usually double and short, sometimes single and 2-fid; alar cells irregular to rectangular, , region clearly differentiated, ± rectangular, long axis parallel to margins; basal laminal cells shorter, longer, or similar to medial cells, ; medial cells short-fusiform to linear-flexuose, 35-60(-64) × (4-)5-6 µm; apical cells shorter; marginal cells similar to medial cells. Sexual condition autoicous; perichaetial inner leaves linear-lanceolate to linear, . Seta yellow-red, orange-red, or red, 0.8-2 cm. Capsule with endostome cilia rudimentary or absent.