Plants soft, yellow-green, sometimes with rusty mottling, or less often yellow-brown, bright green, or blackish green, regularly grading from one color to another within same plant. Stems to 6 cm, usually leafy throughout, irregularly branched; hyalodermis absent, epidermal cells small, walls thick, similar to subadjacent cortical cells, central strand well developed. Leaves usually imbricate, often julaceous, straight or falcate, rarely almost circinate, little different when dry or moist, often oblong-lanceolate, lanceolate, ovate, or occasionally broadly ovate, not deeply concave, (0.5-)1-1.5(-2.5) × (0.3-)0.4-0.8(-1.1) mm; margins regularly involute, not recurved at apex, entire; apex acute or slightly short-apiculate; costa single and short to percurrent, double and short, or ecostate, often within same plant; alar cells many, quadrate, short-rectangular, or irregular, , region usually clearly differentiated; basal laminal cells wider, shorter than medial cells; medial cells short-rhombic to linear-flexuose, (28-)30-55(-95) × (4-)5-7(-12) µm; apical cells shorter; marginal cells rarely longer than 55 µm. Sexual condition autoicous; perichaetial inner leaves lanceolate, . Seta yellow-brown, 0.8-2.4 cm. Capsule with endostome cilia 1-3.