Stems densely rhizoidiferous. Leaf margins occasionally weakly serrulate near apex, occasionally 2-stratose on margins or medial portion of the lamina, marginal cells rhomboid and thick-walled near apex, proximal laminal cells 20-25 µm wide, 2-4:1, walls thin. Interior perichaetial leaves ca. 1.5 mm. Seta red-brown, twisted clockwise proximally, often also counterclockwise distally. Theca red-brown, commonly sulcate, peristome teeth with many articulations, twisted 1.5-2 times, proximal membrane 45-100 µm in height, granulate to spiculose, operculum 0.8-1.5 mm, cells twisted counterclockwise. Calyptra smooth, 1.4-2.1 mm. Spores light brown, weakly papillose.