Plants large, golden to rusty green. Stems 3-15 cm, yellowish brown, creeping to suberect, irregularly to distantly pinnate, branches 2-5+ cm; hyalodermis absent, central strand weakly developed; pseudoparaphyllia foliose, lanceolate to ovate, . Stem leaves falcate-secund, ovate-lanceolate to ovate, gradually narrowed to apex, 0.2-3 × 0.1-1 mm; base not decurrent, not or sometimes slightly auriculate in supra-alar region; margins sometimes recurved proximally, rarely nearly to apex, somewhat toothed in apex; apex acuminate; costa double, faint to distinct; alar cells (usually pigmented in interior, hyaline to yellowish on margins), region well defined, , 3 or 4 cells deep, 5-8 cells wide at base, inner cells grading into basal cells; basal laminal cells shorter, wider than medial cells, sometimes brownish yellow, walls porose, pitted; medial cells 50-80 × 4-6 µm, . Sexual condition dioicous or phyllodioicous. Sporophytes unknown in North America.