Ilex cornuta Lindl. & Paxton

Horned holly (en)

Species

Angiosperms > Aquifoliales > Aquifoliaceae > Ilex

Characteristics

Shrubs or small trees, evergreen, (0.6-)1-4 m tall. Young branchlets longitudinally ridged and sulcate, puberulent or glabrescent along groove; older branchlets gray-white, subterete, longitudinally minutely fissured, leaf scars raised, lenticels absent. Petiole 4-8 mm, adaxially narrowly sulcate and puberulent; leaf blade abaxially greenish, adaxially deep green, shiny, quadrangular-oblong, rarely ovate, 4-9 × 2-4 cm, thickly leathery, both surfaces glabrous, midvein impressed adaxially, lateral veins 5 or 6 pairs, anastomosing near margin, evident abaxially, obscure adaxially, base rounded or subtruncate, margin with 1 or 2 spines per side, sometimes entire, apex with 1 strong spine often reflexed. Inflorescences: cymes, fasciculate, axillary on second year’s branchlets; persistent scales suborbicular, pubescent and ciliate; bracts ovate, pubescent, ciliate, apex obtuse or mucronate; flowers yellowish, 4-merous. Male inflorescences: cymes of order 1 or 2, 1-3-flowered; pedicels 5-6 mm, glabrous; bracteoles basal, broadly deltoid; calyx discoid, ca. 2.5 mm in diam., lobes broadly deltoid, ca. 0.7 × 1.5 mm, membranous, sparsely puberulent, ciliate; corolla rotate, ca. 7 mm in diam., petals reflexed, oblong-ovate, 3-4 mm, basally connate; stamens ca. as long as or slightly longer than petals, anthers oblong-ovoid, ca. 1 mm; rudimentary ovary subglobose, apex obtuse or rounded, indistinctly 4-lobed. Female inflorescences: 1-flowered cymes; pedicels 7-9 mm, glabrous; bracteoles 2, basal, broadly triangular; calyx and corolla as in male flowers; staminodes ca. 4/5 as long as petals, sterile anthers ovoid-sagittate; ovary oblong-ovoid, 3-4 mm, ca. 2 mm in diam., stigma discoid, 4-lobed. Fruit red, globose, 8-10 mm in diam.; fruiting pedicel 1.3-1.4 cm; persistent calyx quadrangular; persistent stigma discoid, distinctly 4-lobed; pyrenes 4, obovoid or ellipsoidal, 7-8 mm, ca. 5 mm in diam., rugose and rugose-pitted throughout, abaxially longitudinally 1-sulcate, endocarp bony. Fl. Apr-May, fr. Aug-Dec. 2n = 38*.
More
An evergreen shrub. It grows to 3.5 m high. It spreads 4.5 m wide. The leaves are thick and glossy. They are almost rectangular and spiny. It is self fertile. The fruit are berries. They are larger than English holly. There are several named varieties. The fruit are round and red and 1 cm across.
Life form perennial
Growth form shrub
Growth support free-standing
Foliage retention evergreen
Sexuality dioecy
Pollination entomogamy
Spread -
Mature width (meter) 4.5
Mature height (meter) 2.4 - 4.0
Root system -
Rooting depth (meter) 0.5
Root diameter (meter) -
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It is a temperate plant. It can tolerate frost. It suits mild-winter climates. In temperate areas in China it grows between 100-1,900 m above sea level. It suits hardiness zones 6-10. Mt Lofty Botanical Gardens.
More
Ravines and hills, 150-600 metres. Shrubby areas, sparse forests, hillsides, streamsides, roadsides, near villages; at elevations from 100-1,900 metres.
Light 6-9
Soil humidity 4-6
Soil texture 2-5
Soil acidity 2-6
Soil nutriment -
Hardiness (USDA) 7-9

Usage

Caution:
Uses environmental use medicinal
Edible fruits
Therapeutic use Abortifacient (unspecified), Arthritis (unspecified), Back (unspecified), Carminative (unspecified), Contraceptive (unspecified), Depurative (unspecified), Fever (unspecified), Intertrigo (unspecified), Lumbago (unspecified), Tonic (unspecified), Tuberculosis (unspecified), Anodyne (unspecified), Refrigerant (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by cuttings or seedlings.
Mode cuttings seedlings
Germination duration (days) 14
Germination temperacture (C°) 22
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -18
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Leaf

Ilex cornuta leaf picture by Remy Way (cc-by-sa)
Ilex cornuta leaf picture by Craig Schneider (cc-by-sa)
Ilex cornuta leaf picture by timothy loftis (cc-by-sa)

Fruit

Ilex cornuta fruit picture by James Thigpen (cc-by-sa)
Ilex cornuta fruit picture by Goldberg Alex (cc-by-sa)
Ilex cornuta fruit picture by nicole macias (cc-by-sa)

Distribution

Ilex cornuta world distribution map, present in China, Korea (Democratic People's Republic of), and United States of America

Conservation status

Ilex cornuta threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:83182-1
WFO ID wfo-0000729319
COL ID 3PFBF
BDTFX ID -
INPN ID -
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Ilex fortunei Ilex furcata Ilex cornuta f. burfordii Ilex cornuta f. gaetana Ilex cornuta var. burfordii Ilex cornuta var. fortunei Ilex cornuta