Annual or perennial herbs or shrubs, not usually strongly scented, lacking latex. Lvs usually alternate, rarely opposite. Involucral bracts in (1)-2-many rows, membranous or foliaceous. Receptacle with or without scales. Capitula usually ☿, rarely unisexual, usually discoid with outer florets ♀ and inner florets ☿, sometimes inner florets ♂ or outer florets ligulate. Anthers sagittate and usually tailed at base. Style branches each with 2 stigmatic lines. Achenes usually homomorphic, rarely heteromorphic, smooth, ribbed or angled, terete or compressed; pappus usually of 1 row of toothed to plumose hairs, sometimes also of scales or rarely 0.