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Trees, monoecious. Leaves in spirals, even-pinnate; leaflet blades glabrous, margin entire. Thyrses axillary or subterminal. Flowers appearing bisexual. Calyx 4-or 5-parted almost to base, imbricate. Petals 4 or 5, distinct, contorted, much longer than calyx when in bud. Staminal tube urceolate, kettlelike, or cup-shaped; anthers 8-10, inserted on inside apical part of tube. Disk cushion-shaped. Ovary 4-or 5-locular, with 12-16(-18) amphitropous ovules per locule; stigma disciform, with 4 grooves on apex. Capsule globose or suborbicular, woody, apically 4-or 5-valvate when mature. Seeds with vestigial endosperm; cotyledons flat, collateral; radicle lateral.
Ovary 4–5-locular, each loculus with 12–16 or more pendulous ovules in 2 rows; style short; style-head discoid, almost completely blocking entrance to staminal tube; margin crenellate, upper surface with minute central papilla with 4–5 radiating, (?) stigmatic ridges.
Staminal tube urceolate, glabrous inside and out, bearing 8 or 10 included anthers or antherodes inside towards the apex and terminated by 8 or 10 subcircular, emarginate or irregularly lobed appendages alternating with the anthers or antherodes.
Fruit an upright, almost spherical woody septifragal capsule, opening by 4–5 valves from the apex; columella not extending to apex of capsule, with 4–5 sharp ridges, hard and woody; seed scars white, conspicuous.
Flowers monoecious, but with well-developed vestiges of opposite sex, and with little external difference between the sexes, borne in large much-branched panicles.
Seeds 6 or more per loculus, broadly transversely ellipsoid, narrowly winged more or less all round margin.
Pistillode similar to ovary but much narrower; loculi well-developed, but vestigial ovules minute.
Disk cushion-shaped, fused to the base of the ovary but free from the staminal tube.
Petals 4–5, free, much longer than the calyx in bud, contorted.
Calyx lobed almost to base, lobes 4–5, subcircular, imbricate.
Leaves paripinnate; leaflets entire.
Trees, usually of large size.
Pending.
