Annual or perennial herbs or subshrubs, unarmed. Raphides present. Leaves opposite or sometimes ternate, without domatia; stipules persistent or deciduous, interpetiolar and fused to petioles and/or shortly united around stem, 2-6-lobed or-setose, with apices of segments or setae usually glandular. Inflorescences terminal, thyrsiform, subcapitate, cymose, or often corymbiform with axes sometimes elongating and becoming racemiform with age, several to many flowered, pedunculate or sessile, bracteate. Flowers sessile or pedicellate, bisexual, distylous. Calyx limb 4-lobed; lobes sometimes unequal. Corolla white, pink, lilac, or violet, funnelform, salverform, or tubular, with shape sometimes differing between long-styled and short-styled forms, lanate inside tube; lobes 4, valvate in bud. Stamens 4, inserted in corolla throat or near middle of corolla tube, included or exserted; filaments short; anthers dorsifixed. Ovary 2-celled, ovules 1 in each cell, pendulous, apical; stigma 2-lobed, exserted or included. Fruit schizocarpous, ovoid to ellipsoid, sometimes laterally compressed and/or didymous, dry, with calyx limb persistent; mericarps 2, ellipsoid, indehiscent, with 1 seed, early to tardily separating from base upward and falling together with or separately from a carpophore, with carpophore variously filiform and basal to comprising entire enlarged septum; seeds medium-sized, oblong-ellipsoid, compressed; testa thin; endosperm fleshy; cotyledons thin; radicle ascending.
Home