Plants small to large, pale to dark green or golden. Stems terete-foliate, irregularly pinnate to regularly 2-or 3-pinnate, often somewhat stipitate; hyalodermis absent, cortical cells small, walls incrassate; paraphyllia absent; pseudoparaphyllia foliose; axillary hair basal cells 1 or 2, short, brown, apical cells 1-4, elongate, hyaline. Stem and branch leaves often differentiated, loosely appressed, usually ovate, usually concave; base decurrent; costa single or double, short, sometimes weak on stipe and stem leaves and strong on branch leaves; alar cells usually well differentiated, usually quadrate, often excavate; medial laminal cells short to linear, smooth or prorulose. Sexual condition dioicous. Seta long or sometimes short, smooth or roughened throughout or distally. Capsule erect or sometimes horizontal, cylindric, symmetric or sometimes curved; annulus usually differentiated; operculum conic to short-rostrate, straight; peristome double, perfect or sometimes reduced; exostome, when not reduced, with external surface cross striolate proximally, papillose distally, when reduced, striate, papillose, or smooth; endostome basal membrane high or low, segments long, narrowly perforate, cilia nodose or appendiculate. Calyptra cucullate, smooth or slightly roughened, naked or hairy.
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