Trees', bark greyish brown, slightly fissured and/or with lenticels; indumentum of simple hairs only. Branchlets smooth to slightly grooved, sericeous when young. Leaves paripin-nate, pulvinate at base, no pseudo-stipules or wings; petiolules present as pulvinus. Leaflets subopposite to opposite, basal ones ovate, upper ones elliptic (to obovate), slightly asymmetric, acroscopic side broader, thin, not punctate; base usually cuneate, asymmetric; margin entire, flat; both surfaces smooth, subglabrous except for at least the subseri-ceous basal part of the midrib, domatia as (a few) glabrescent hair tufts below; venation raised, especially below, nerves marginally looped, veins reticulate. Inflorescences ram-iflorous to axillary to pseudoterminal, reduced thyrses with usually 3 cymules per node; rachis (sub)sericeous; cymules basally dichasial to apically cincinnate, those with male flowers often with an exceptional number of sepals, petals, and stamens in first developed flowers. Bracts and bracteoles: mainly margin and outside sericeous. Pedicels with glandular hairs besides simple ones, articulated. Flowers actinomorphic, yellowish white. Sepals (sub)equal, basally united; tube disc-shaped, c. 0.3 mm long; blades triangular (to ovate). Petals: claw c. 0.1 mm high; blade triangular; scales united into one, larger than blade, margin irregularly crenate, pilose; crests usually absent (to present). Disc circular, smooth, glabrous, enveloping base of stamens, without appendages. Stamens (7-)8(-10); filaments especially basally pilose; anthers basifixed, dehiscence latrorse, pilose. Pistil: ovary flat, 2-(or 3-)locular; ovules one per locule; style and stigma united, flat, triangular, stigmatic part folded outwards like overhanging roof. Fruit an obovoid, loculicidally dehiscing capsule, with usually 1 developed seed, flat when 2-locular, smooth to somewhat rough, outside glabrous, margin sharp, almost slightly winged, inside glabrous to pilose, red; stipitate or not; wall thin, coriaceous to woody, at most 2 mm thick. Seeds ellipsoid, triangular in transverse view, base straight to oblique, smooth, shiny, black; sarcotesta glabrous, covering seed only basally to almost completely, orange; testa thin, without radicle pocket on the inside.
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Monoecious trees. Branchlets lenticellate, usually hairy towards apex, with simple hairs. Leaves paripinnate; leaflets opposite or alternate, usually decreasing in size from apex, entire. Inflorescence axillary, panicle-like. Flowers regular. Calyx cupular, 5-partite; lobes ovate, valvate. Petals 5, minute; scale broad, free or united to margin forming a broad funnel or 2-valved and broadly ovate; crests absent. Disc complete. Stamens 8–10; filaments and anthers hairy. Ovary sessile, compressed, 2-locular; ovule 1 per locule, 1 usually abortive; style short. Fruit obovoid or obcordate, 2-sulcate, turgid but slightly compressed at sutures, 1-seeded, 2-valved; valves thick, drying crustaceous or woody. Seed subglobose; aril fleshy, basal or nearly covering seed.
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