Plants medium-sized, sometimes smaller and slender, grayish green to yellowish green, dull yellowish to pale brown or yellowish brown, not glossy, in compact or loosely interwoven mats; leafy stems julaceous, mostly ca. 2-4 cm long, (0.30-)0.45-0.65(-0.80) mm wide, irregularly branched or subpinnately branched, sometimes bearing many young short, julaceous branchlets, usually curved at tips, sometimes clavate (from wet habitat); branchlets 2.5-3.5 mm long, sometimes filiform branches present; stems in cross section with 2-3 layers of small, thick-walled epidermal cells, a cortex of enlarged, hyaline, thin-walled cells, central strand weakly developed; pseudoparaphyllia foliose. Main stem leaves 1.0-1.3 x 0.5-0.7 mm, imbricate when dry, erect-tumid when wet, broadly ovate at base, somewhat concave and more or less symmetric, reflexed or secund to subsecund at apex, abruptly short-acuminate, ending in a point 1-2 cells long; margins entire below, often clearly serrate; costae thin, double or often forked above the base, reaching up to 1/3-1/2 the leaf length, in cross section cells not very differentiated, stereids absent; upper leaf cells rhombic to short rhomboidal, 16-19 x 8-9 µm; median cells rhomboidal, 28-32 x 7-8 µm, verrucose, moderately thick-walled; basal juxtacostal cells rounded-quadrate to rounded-rhombic, ca. 7-9µm wide; alar cells quadrate or rounded-quadrate, in 6-9 rows toward costa, forming a distinct alar group; branch leaves similar to stem leaves, but smaller. Dioicous. Perichaetial leaves oblong-lanceolate, singly costate. Setae 0.9-1.1 cm long, smooth; capsules erect, oblong-ovoid to short-cylindrical, 2.2-2.5 x 1.0-1.2 mm; opercula conic, 0.35-0.45 mm high; exothecial cells short-rectangular or irregularly rounded-pentagonal, thick-walled, not collenchymatous; stomata immersed; annuli in 3-4 rows of small, quadrate, thick-walled cells; peristome double, exostome teeth dark brown, lanceolate, inserted well below the mouth, 0.35-0.45 mm high, papillose throughout, outer surface with zig-zag line, inner surface strongly trabeculate; endostome segments shorter than the teeth, 0.20-0.25 mm high, basal membrane 70-90 µm high; cilia absent. Spores16-19 µm in diameter, papillose. Calyptrae not seen.
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