Stems spreading and recurved distally. Stem leaves on young shoots and subtending branches green to pale green, those below branches reddish brown proximally and hyaline distally, slightly contorted and rarely somewhat crisped when dry, appressed to stem proximally and slightly spreading distally, ± flat when moist, (2-)2.5-3(-4) mm; margins hyaline or reddish brown, teeth large, more pronounced distally, often at right angles to margin; apex long-acuminate; medial laminal cells elongate, 20-50 µm; marginal cells linear and short-linear, in 1 or 2 rows. Branch leaves green to dark green, moderately crisped when dry, spreading, keeled when moist, 1.3-2(-2.5) mm; margins green, teeth small, more pronounced distally, not at right angles to margin; apex acute, cuspidate; medial laminal cells short-elongate, short-rhombic, or ± isodiametric, 17-27µm marginal cells short-linear and rhomboidal, in 1 (or 2) rows. Capsule with annulus differentiated in 3 rows of cells; exostome papillose; endostome papillose, basal membrane porous, cilia 2 or 3.
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