Lindsaea javanensis Blume

Rhizome short-creeping, 1-2 mm ø; scales medium brown, lanceolate, long-acuminate, to 1.5 mm long, to 4-seriate at base, with a long uniseriate apex. Leaves clustered; petioles 10-30 cm long, especially in large leaves much longer than the lamina, dark reddish brown to atropurpureous, ± lustrous, abaxially terete at base, upward gradually obtusely, at apex mostly acutely bi-angular, the angles and especially the borders of the adaxial groove often pale. Lamina herbaceous to chartaceous, dark green or olivaceous when dry, very variable, triangular or oblong or in small leaves sometimes transvers ely triangular, 7-20 cm long, 6-15 cm wide, 1.25 to (in large leaves) 1.75 times as long as wide, simply pinnate to amply bipinnate, in the first case with up to as little as 2 pairs of lateral pinnules and a distinct but not conform terminal one, in the second case with up to 5 (pinnate) pinnae to a side and several simple ones above, which are gradually reduced to and confluent (or not) with the terminal pinnule (segment). Primary rachis dark, adaxially with a narrow, pale-edged groove, abaxially flattened, pale-angled. Primary pinnae their width apart to contiguous, spreading or in plurijugate bipinnate leaves ascending. Pinnules of simply pinnate or subbipinnate leaves rhombic, very often with long-acuminate to caudate apex, with very unequal base, basiscopically much more cut away, to c. 8 by 2 cm; apex of lamina similar but ± symmetric at base; apices of fully pinnate pinnae similar but smaller, progressively smaller and less pointed as there are more secondary pinnules; larger transitional pinnules between pinnate pinnae and the lamina apex often rhombic-caudate, smaller ones subtrapeziform or subflabellate, very obtuse; pinnules of fully pinnate pinnae up to 8 to a side, mostly not contiguous, subtrapeziform, rounded-rhombic or subflabellate. Secondary rachises abaxially flattened or slightly sulcate, abruptly pale at their insertion on the dark primary rachis, usually distinctly green-margined. Fertile pinnule-margin subentire or minutely erose, in larger leaf segments here and there incised by shallow crenations; sterile margin, especially near the segment bases, sharply serrate or dentate, with deeper incisions. Lobes of pointed pinnules often slightly concave. Veins immersed but +-evident, free, mostly twice forked, c. 1 mm apart; larger pointed pinnules and terminal divisions with a percurrent costa. Sori continuous in small pinnules, progressively more interrupted in larger ones, bi-to plurinerval. Indusium pale, subentire to erose, 0.3-0.4 mm wide, almost reaching the margin to falling short of it by more than its width, little reflexed at maturity. Spores light brown, trilete, smooth, c. 25 μ.

Terrestrial in forests, 80-1400 m, mostly above 800 m.