Trees, monoecious. Indumentum of solitary, simple or 2-branched hairs; no glandular scales. Leaves spirally arranged or sometimes partly, nearly opposite, especially towards the inflorescence, paripinnate, 1-4-(?sometimes 5-)jugate; pseudo-stipules absent; petiole and rachis not winged. Leaflets mostly (sub)opposite, beneath dull, margin entire to undulate, mainly in the upper part. Inflorescences terminal and axillary, thyrsoid; bracts triangular, 0.5-2 mm long. Flowers actinomorphic, unisexual. Calyx 4-or 5-merous, outside and inside densely appressed short-hairy, the tube cupular, the lobes to 0.3-0.5 of the length of the tube, apert in bud, equal, not petaloid. Petals absent. Disc annular, small, without appendages. Stamens 6-11, far exserted in male flowers, short in female and in seemingly bisexual, but functionally male flowers; filaments variably hairy; anthers glabrous. Pistil short-stalked; ovary 2-(?or 3-)celled; style terminal, shorter than the ovary; stigma consisting of 2 (or 3?) long, spreading to recoiled lobes. Ovules 1 per cell, basal. Fruits nearly always only 1 cell developed, hardly stipitate, ellipsoid, ovoid, or globular, not winged, indehiscent(?), glabrous, spiny to scaly; pericarp thin, coriaceous to rather hard, endocarp glabrous. Seeds glabrous, partly or completely enveloped by an entire, translucent, juicy arillode.
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Trees, monoecious. Leaves paripinnate, alternate, estipulate. Thyrses terminal, golden tomentose; bracts and bracteoles small. Flowers unisexual, actinomorphic. Calyx cupular, 4-or 5-lobed, valvate, opening early. Petals absent. Disk acetabuliform, entire. Stamens (male flowers) 6-8, exserted; filaments filiform, pilose. Ovary (female flowers) shortly stalked, obcordiform, 2(or 3)-lobed, 2(or 3)-loculed; ovules 1 per locule; style inserted between ovary lobes; stigma 2-or 3-lobed. Fruit deeply parted into 2 or 3 schizocarps, usually only 1 or 2 developed, ovoid or subglobose; pericarp leathery (crustaceous when dry), abaxially with tortoise-shell-like fissure, with scattered conical prominents, sometimes nearly smooth. Seeds nearly same shape as schizocarps; testa brown, nitid, leathery, arillode fleshy, wrapping all or lower half of seed; embryo erect. 2n = 28, 30.