Bathyphylls with up to 28 pairs pinnae have lobed terminal lamina continuous with rachis; lower pinnae gradually reduced, lowest c. 7 by 3 mm, largest pinna 2½ by 0.7 cm, base unequally cuneate (more narrowly on basiscopic side), edges crenulate with teeth 2-3 mm apart, apex often not acute, bases and apices of lower pinnae more rounded; stipe, rachis and costae rather densely scaly, scales dark; pinnae reddish when dry; veins strongly raised on both surfaces. Sterile acrophylls: stipes 20-30 cm, pinnae commonly 1.2-1.8 cm wide, 12-15 cm long (largest seen 22 by 2.2 cm), base slightly unequally subtruncate to subcordate, edges subentire and parallel, apex acuminate and somewhat falcate, irregularly sinuate or somewhat toothed, uppermost pinnae gradually smaller with less truncate base and more toothed apex, lowest pinnae not much reduced but often with cordate base; texture thin, colour often reddish when dry; veins slightly prominent, areoles rather small, dark scales rather abundant. Fertile fronds: pinnae commonly to 20 cm long, base unequally cuneate, widening suddenly from a winged stalk to 10 mm long; base of pinna 7-10 mm wide, only the edges fertile, upper part gradually narrowed, distal half sometimes covered with sporangia but often with a narrow sterile band each side of midrib.
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