Bathyphylls: stipes 3-10 cm long; lamina to 40 by 9 cm; pinnae 20-30 pairs, lowest rather reduced and deflexed, upper gradually smaller, uppermost grading into lobes of frond-apex which is not pinna-like not jointed; middle pinnae to 5 by 1.2 cm, at right angles to rachis, acroscopic base broadly subtruncate to subauriculate, basiscopic narrower, rounded, sides almost parallel for ⅔ of length, then tapering to acute apex; edges shallowly crenate with 1-2 teeth at distal end of each crena-tion; texture thin, veins slender, prominent. Sterile acrophylls: stipes to 20 cm, frond to at least 90 cm long; lowest pinnae somewhat reduced and more distant, uppermost gradually smaller; middle pinnae to 18 by 2.5 cm, sessile or subsessile, acroscopic base broadly cuneate, basiscopic narrower and rounded to cuneate; edges usually with a few irregular teeth towards acuminate apex; texture thin, veins conspicuous. Fertile pinnae to 15 cm by 5 mm, on stalks to 2 mm, upper ones gradually reduced, uppermost c. 4 cm long.