Lycopodium clavatum L.

Stag's-horn clubmoss (en), Lycopode en massue (fr), Éguaire (fr)

Species

Pteridophytes > Lycopodiales > Lycopodiaceae > Lycopodioideae > Lycopodium

Characteristics

Plants terrestrial. Stems wide-creeping, rooting at irregular intervals, anisotomously branched, main stems with indeterminate growth, to 2 mm in diameter, isophyllous, lateral branches erect, arising in a dorsi lateral position on the main stem, isophyllous, fertile branches erect, anisotomously branched, sparsely leafy, isophyllous, heterophyllous, to 180 mm long. Leaves herbaceous, acicular, entire, aristate, patent to imbricate, to 7 mm long, to 1 mm wide. Strobili terminal, in groups of 3-5, to 30 mm long, isophyllous, sporophylls ephemeral, herbaceous, subpeltate, often with a basiscopic membranous wing on the stalk, narrowly to broadly ovate, aristate, laciniate, to 4.5 mm long, to 2 mm wide. Sporangium reniform, short-stalked, borne adaxially near the sporophyll base, isovalvate, to 1.3 mm in diameter. Spores spheroidal to subtriangular, trilete, reticulate, (34-)37.5(-38) µm in equatorial diameter, (30-)31 (-34) µm in polar diameter.
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Stolons subterranean, slender and creeping, 1 or 2 times forked, green, with sparse entire leaves; lateral branches erect, 20-25 cm tall, 3-5 times forked, sparse, flattened (young branches terete), stem together with leaves 9-12 mm in diam. Leaves spirally arranged, dense, angled upward, lanceolate, 4-6 × ca. 1 mm, leathery, with transparent hairs, midrib visible on both surfaces, base cuneate, decurrent, sessile, margin entire, apex acuminate. Strobili 2(or 3) together on a peduncle, peduncle up to 12 cm, bracts of peduncle spirally arranged, sparse, herbaceous, narrowly lanceolate; strobili erect, terete, 3.5-4.5 cm × ca. 4 mm, subsessile or with short, equally long pedicels; sporophyll broadly ovate, ca. 1.5 × 1.3 mm, papery, margin membranous, erose, apex acute, with short acute tip. Sporangia slightly exposed.
Horizontal stems superficial, elongate, arching and rooting at intervals, 2–3 mm thick, not dorsiventral, the lvs curved upwards; erect stems branched, to 2.5 dm; lvs mostly 10-ranked, uniform, linear-subulate, 4–7 × 0.5–0.8 mm, slightly or scarcely toothed, attenuate to a hyaline hair-like tip 2–3 mm; cones 1.5–11 cm, 3–7 mm thick, 1–6 on remotely bracteate peduncles 1.5–15 cm; sporophylls yellowish, broad-based, abruptly acuminate, with scarious erose margins; sporangia reniform, ca 1.5 mm wide; spores 28–36 microns in diameter, the sides convex, the commissural and outer faces angular-reticulate, the commissures not in furrows; 2n=68. Open dry woods and rocky places in acid soil; circumboreal, s. in e. U.S. to N.C., W.Va., Mich., and Io.
Horizontal stems on substrate surface. Upright shoots clustered, 0.6--1.2 cm diam., dominant main shoot with 3--6 branches mostly in lower 1/2. Lateral branchlets few and like upright shoots; annual bud constrictions abrupt, branchlets mostly spreading. Leaves spreading, often somewhat ascending in distal 1/3 of branches, medium green, linear, 4--6 X 0.4--0.8 mm; margins entire; apex with narrow hair tip 2.5--4 mm. Peduncles 3.5--12.5 cm, with remote pseudowhorls of appressed leaves, loosely branched into 2--5 alternate stalks, 0.5--0.8 cm. Strobili 2--5 on alternate stalks (if double, usually with stalks 5--8 mm), 15--25 X 3--6 mm. Sporophylls 1.5--2.5 mm, apex abruptly reduced to hair tip. 2 n = 68.
Terrestrial. Main stem creeping, up to 3 mm in diameter, producing dichotomously branched upright stems at 50-100 mm intervals. Leaves narrowly lanceolate, patent to suberect or imbricate, up to 5 x 0.5 mm, with a translucent hair point up to 3 mm long apically, margin usually entire but sometimes ciliate in leaves on main stem. Strobili pedunculate, 20-80 mm long in groups of 2-5 (rarely solitary) at apex of sparsely leafy peduncle up to 190 mm long. Sporophylls broadly ovate, acuminate, 2-3 x 1.5-2 mm, margin finely lacerate.
Stems widely creeping, irregularly branched, up to 2 mm in diam., lateral branches erect, up to 180 mm long, sparsely leafy. Lycophylls herbaceous, acicular, with a translucent hair point, entire, up to 7 x 1 mm. Strobili pedunculate, in groups of 2-5, up to 30 mm long. Sporophylls ephemeral, herbaceous, narrowly to broadly ovate, laciniate, up to 4.5 x 2 mm. Sporangium reniform, up to 1.3 mm in diam.
Terrestrial. Stems erect, branching from a prostrate, branched main stem, 80-300 mm long. Leaves erect from branches at right angles, with aristate tips bent forward, 3-5 mm long, up to 0.6 mm wide; sporophylls ephemeral, distinct from sterile leaves. Sporangia short-stalked, borne in sporophyll axils in a terminal strobilus.
Sporophylls different from foliage leaves, grouped into pedunculate strobili. Horizontal stem producing much-branched, erect stems at intervals. Leaves hair-pointed. Erect stems not laterally compressed.
Perennial with prostrate main stem and dichotomously branched, erect stems. Leaves with a translucent hair point. Strobili pedunculate, in groups of 2-5.
A herb. It grows 40 cm tall.
Life form perennial
Growth form herb
Growth support -
Foliage retention evergreen
Sexuality monoecy
Pollination hydrogamy
Spread anemochory
Mature width (meter) 1.0
Mature height (meter) 0.1
Root system creeping-root
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

Moorland, fields and pastures. Cool, wet climates where it prefers open habitats such as mountain sides, moors and heaths, clearings and road cuttings in cloud forest. In the tropics it grows in highland areas above 1,300 metres.
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It is a temperate plant. In northern China it grows in conifer and moss forests between 700-1,800 m above sea level. In Argentina it grows between 600-2,500 m above sea level. It suits hardiness zones 1-8.
Moorland, fields and pastures, it is rare in lowland areas.
Light 1-8
Soil humidity 4-6
Soil texture 5-8
Soil acidity 2-7
Soil nutriment -
Hardiness (USDA) 6-11

Usage

The leaves are used for a herbal tea.
Uses animal food dye environmental use fiber food fuel material medicinal oil social use tea
Edible leaves
Therapeutic use Hemostat (fruit), Diarrhea (reproductive organ), Dyspepsia (reproductive organ), Emetics (reproductive organ), Erectile dysfunction (reproductive organ), Exanthema (reproductive organ), Kidney calculi (reproductive organ), Liver cirrhosis (reproductive organ), Liver diseases (reproductive organ), Postpartum hemorrhage (reproductive organ), Scurvy (reproductive organ), Skin diseases (reproductive organ), Urination disorders (reproductive organ), Wounds and injuries (reproductive organ), Fatigue (shoot), Myalgia (shoot), Analgesic (unspecified), Gynecological Aid (unspecified), Febrifuge (unspecified), Stimulant (unspecified), Headache (unspecified), Apertif (unspecified), Cancer(Skin) (unspecified), Carminative (unspecified), Catarrh (unspecified), Chest (unspecified), Diarrhea (unspecified), Diuretic (unspecified), Dyspepsia (unspecified), Emmenagogue (unspecified), Fever (unspecified), Fumitory (unspecified), Homeopathy (unspecified), Laxative (unspecified), Nerves (unspecified), Pressor (unspecified), Preventitive(Chafe) (unspecified), Rheumatism (unspecified), Skin (unspecified), Spasm (unspecified), Uterotonic (unspecified), Aphrodisiac (unspecified), Urogenital (unspecified), Anti-infective agents, local (unspecified), Antirheumatic agents (unspecified), Asthenia (unspecified), Constipation (unspecified), Demulcents (unspecified), Digestive system diseases (unspecified), Diuretics (unspecified), Eczema (unspecified), Emollients (unspecified), Epilepsy (unspecified), Gastritis (unspecified), Herpesvirus 4, human (unspecified), Hypnotics and sedatives (unspecified), Hypoglycemic agents (unspecified), Kidney diseases (unspecified), Liver cirrhosis (unspecified), Liver diseases (unspecified), Lung diseases (unspecified), Menstruation-inducing agents (unspecified), Mycoses (unspecified), Parasympatholytics (unspecified), Urinary bladder, overactive (unspecified), Urination disorders (unspecified), Urologic diseases (unspecified), Nocturnal enuresis (unspecified), Antirheumatic agents (whole plant), Diuretics (whole plant), Kidney diseases (whole plant), Parasympatholytics (whole plant)
Human toxicity toxic (whole)
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Lycopodium clavatum habit picture by Юрий Басов (cc-by-sa)
Lycopodium clavatum habit picture by Yoan MARTIN (cc-by-sa)
Lycopodium clavatum habit picture by Yoan MARTIN (cc-by-sa)

Leaf

Lycopodium clavatum leaf picture by gisèle Friso (cc-by-sa)
Lycopodium clavatum leaf picture by Jesse Corriveau (cc-by-sa)
Lycopodium clavatum leaf picture by Jakub Jan (cc-by-sa)

Flower

Lycopodium clavatum flower picture by Nicolas Roux (cc-by-sa)
Lycopodium clavatum flower picture by Lars Bjerges (cc-by-sa)
Lycopodium clavatum flower picture by hieracium eriophorum (cc-by-sa)

Fruit

Lycopodium clavatum fruit picture by Yoan MARTIN (cc-by-sa)
Lycopodium clavatum fruit picture by Yoan MARTIN (cc-by-sa)
Lycopodium clavatum fruit picture by huy HO (cc-by-sa)

Distribution

Lycopodium clavatum world distribution map, present in Angola, Andorra, Argentina, Austria, Burundi, Belgium, Bulgaria, Bosnia and Herzegovina, Belarus, Bolivia (Plurinational State of), Brazil, Canada, Switzerland, China, Côte d'Ivoire, Cameroon, Congo, Colombia, Comoros, Costa Rica, Cuba, Czech Republic, Germany, Denmark, Ecuador, Spain, Estonia, Ethiopia, Finland, France, Gabon, United Kingdom of Great Britain and Northern Ireland, Georgia, Guinea, Guadeloupe, Equatorial Guinea, Guatemala, Guyana, Honduras, Croatia, Hungary, Iceland, Italy, Jamaica, Japan, Kazakhstan, Kenya, Liberia, Liechtenstein, Lesotho, Lithuania, Luxembourg, Latvia, Madagascar, Mexico, North Macedonia, Montenegro, Mongolia, Mozambique, Martinique, Mauritius, Malawi, Nigeria, Nicaragua, Netherlands, Norway, Panama, Peru, Poland, Puerto Rico, Korea (Democratic People's Republic of), Portugal, Paraguay, Réunion, Romania, Russian Federation, Rwanda, Sudan, Sierra Leone, El Salvador, Saint Pierre and Miquelon, Serbia, Sao Tome and Principe, Slovakia, Slovenia, Sweden, eSwatini, Thailand, Turkey, Tanzania, United Republic of, Uganda, Ukraine, Uruguay, United States of America, Venezuela (Bolivarian Republic of), South Africa, Zambia, and Zimbabwe

Identifiers

LSID urn:lsid:ipni.org:names:304307-2
WFO ID wfo-0001107341
COL ID 3WMCR
BDTFX ID 40485
INPN ID 107003
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Lycopodium clavatum f. frondosum Lepidotis clavata Lycopodium aristatum Lycopodium piliferum Lycopodium trichophyllum Lycopodium eriostachys Lycopodium mayoris Lycopodium clavatum f. clavatum Lycopodium ciliatum Lycopodium contiguum Lycopodium preslii Lycopodium herbaceum Lycopodium clavatum f. brevipedunculatum Lycopodium clavatum f. decapitatum Lycopodium clavatum f. sterilis Lycopodium trichiatum Urostachys plutonis Lepidotis ciliata Lepidotis inflexa Lycopodium inflexum Lycopodium clavatum subsp. clavatum Lycopodium clavatum var. borbonicum Lycopodium clavatum var. trichiatum Lycopodium clavatum var. inflexum Lycopodium clavatum var. tristachyum Lycopodium clavatum var. clavatum Lycopodium aristatum var. robustius Lycopodium aristatum var. incurvum Lycopodium clavatum var. raddianum Lycopodium clavatum var. aristatum Lycopodium clavatum var. desvauxianum Lycopodium clavatum var. jamaicense Lycopodium aristatum var. desvauxianum Lycopodium trichiatum var. desvauxianum Lycopodium clavatum var. minarum Lycopodium clavatum var. eriostachys Lycopodium clavatum var. piliferum Lycopodium clavatum var. equisetoides Lycopodium clavatum var. trichophyllum Lycopodium clavatum subsp. contiguum Lycopodium clavatum var. pseudocontiguum Lycopodium clavatum var. preslianum Lycopodium vestitum var. herbaceum Lycopodium clavatum var. nataliense Lycopodium clavatum var. robustius Lycopodium clavatum f. robustinus Lycopodium clavatum var. laurentianum Lycopodium clavatum var. montanum Lycopodium clavatum var. paucidivisum Lycopodium clavatum var. serpens Lycopodium clavatum var. sitchaense Lycopodium clavatum var. subremotum Lycopodium clavatum var. asiaticum Lycopodium clavatum