Scandent rachis about 1.5 mm diameter; primary rachis-branches very short, hairs on the dormant apex having swollen bases; sterile secondary rachis-branches once or twice equally or sub-equally dichotomous (12-20 mm to first dichotomy), the leaflets simple or very deeply bilobed, 10-20 cm long, 1.5-2.5 cm wide, the outer base of each more or less cordate (rarely strongly auriculate), edges irregularly serrate and slightly thickened; fertile secondary branches usually sub-pinnate with flexuous axis bearing two lateral and one terminal leaflets, sometimes twice symmetrically dichotomous; fertile leaflets usually deeply bilobed or geminate, one or both members often having a shorter lobe commonly 9-15 cm long and up to 13 mm wide including sorophores, lamina reduced to a wing along the costa and along each vein-group (sometimes two adjacent vein-groups with a common lamina) the sorophores thus in groups of 2-5; sorophores usually 2-3 mm long; spores coarsely and unevenly verrucose, the warts often confluent.
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