Rhizome and juvenile fronds not seen. Rachis of scandent frond to 2½ mm diameter, minutely hairy (hairs very slender, erect); primary rachis-branches very short, the sunken dormant apex bearing pale hairs; secondary rachis-branches also very short (1-3 mm long), so that the leaflets appear to be verticillate on the main rachis; leaflets 3 or 4 on each side of a dormant apex; sterile leaflets c. 20 cm long, 2-2½ cm wide, entire (margin more or less thickened and car-tilaginous), gradually narrowed to the narrow truncate base, sometimes with one or two minute auricles forming separate small lateral leaflets, lamina covered with a close network of conspicuous anastomosing veins, with about 5 rows of areoles between costa and margin (outer areoles progressively smaller), ultimate free veins joining the thickened margin, surface glabrous with scattered warts when dry; fertile leaflets often longer than sterile (to 30 cm long) but narrower, usually reduced to a narrowly winged costa with a row of sorophores on each side, less commonly to a lamina 8 mm wide in which there is slight anasto-mosis of veins; sorophores 3-4 mm long, contracted at base; indusia more or less hairy; spores coarsely verrucose.