Plants yellow-green or olivaceous above, brown below; corticolous. Primary stem up to 50 mm long, branching irregular; secondary stem in section with inner cortex 5 or 6 cells across, incrassate, outer cortical cells in 1 or 2 rows, consisting of substereids or stereids, outer surface rough. Stem leaves crowded to somewhat distant, frequently reflexed; ovate-acuminate, lanceolate or lanceolate-ligulate, 0.8-1.1 mm long. Branch leaves crowded, ± equal in size, appressed to spirally appressed or erect when dry, occasionally twisted, erect-spreading to spreading when wet; occasionally reflexed; lanceolate or oblong, 0.8-1.8 mm long, ± keeled, unistratose, occasionally with bistratose patches; apex acuminate, acute, rounded acute to obtuse, rarely cucullate; margins plane above, plane or recurved below, entire. Costa ending below apex to percurrent or rarely mucronate; ventral superficial cells linear, smooth; dorsal superficial cells oval above, linear below, smooth to bulging; in section crescent-shaped to subround, flat ventrally, bulging dorsally, guide cells absent, 2-4 ventral substereids or stereids present, dorsal surface cells incrassate. Upper laminal cells oval above, oval to rounded quadrate below, incrassate, flat to bulging above, bulging or mammillose below, occasionally with intercellular spaces, 7.5-14.0 µm; basal cells frequently longer and narrower towards costa. Dioicous. Perigonia terminal on branches or branchlets, leaves broadly ovate to ovate-acuminate. Perichaetia terminal on branches or branchlets; leaves oblong, oblong-acuminate, oblong-lanceolate or oblong-ligulate, 1.0-1.8 mm long. Seta 3.2-8.0 mm long, yellowish or reddish to brown, ochrea weakly developed or absent. Capsule elliptic or oblong-cylindrical; urn 0.8-1.4 mm long, yellowish brown to brown, ribbed; neck 0.2-0.8 mm long; exothecial cells irregularly rectangular to quadrate, smaller at mouth, incrassate. Peristome double; outer layer 7-10 cells high, irregularly split, striolate-papillose, yellowish brown; inner membrane frequently plicate, outer surface smooth, inner surface coarsely papillose, hyaline. Operculum 0.5-0.8 mm long. Calyptra 1.5-1.8 mm long, naked or with a few hairs, yellow below, red-brown above. Spores 23-28 µm.
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