Plants medium-sized to large, forming mats, yellowish green, green or greenish brown above, greenish brown to brown below; corticolous. Primary stem up to 100 mm long, branching regularly, tomentose below; rhizoids smooth, reddish brown; secondary stem 2-10 mm tall, branching by subperichaetial innovations; in section inner cortex 10-15 cells across, incrassate, outer cortical cells smaller, in 1-3 rows, consisting of substereids to stereids, outer surface rough. Stem leaves crowded to ± distant, fragile on old stems; ovate-acuminate or ovate-subulate, 1.0-2.3 mm long. Branch leaves crowded, erect, curved to twisted, inrolled when dry, erect-spreading, inflexed when wet, ± rugose, irregularly unistratose to multistratose; narrowly ovate-lanceolate, narrowly lanceolate or narrowly lanceolate-subulate, 1.7-3.0 mm long, keeled; apex acuminate, fragile; margins plane, entire to crenulate-papillose above, entire below. Costa ending below apex to percurrent, ventral and dorsal superficial cells linear, smooth; in section subround to round, bulging dorsally, guide cells frequently differentiated, incrassate, dorsal stereids present. Upper laminal cells rounded, incrassate, bulging, 8.7-15.0 µm, multipapillose, papillae small, low; basal cells rectangular, longitudinal walls incrassate, straight to sinuate, frequently unipapillose or tuberculate. Pseudautoicous; dwarf male plants on secondary stem, leaves ovate or ovate-apiculate. Perichaetia terminal, overgrown by subperichaetial innovations; leaves oblong-lanceolate, 1.6-2.7 mm long, unistratose. Seta 3.5-6.0 mm long, reddish brown. Capsule reddish brown, mouth darker, smooth to weakly ribbed above; urn ovate-cylindrical, 0.8-1.4 mm long; neck up to 0.4 mm long, wrinkled dry; exothecial cells irregularly quadrate to rectangular, smaller at mouth, incrassate. Peristome single; exostome teeth 16, narrowly oblong, blunt, frequently irregular in outline, (125-)150-225 µm long, pale yellow or yellowish brown, hyaline above, erect to incurved dry, striolate-papillose. Operculum 0.8-1.0 mm long. Calyptra 2.8-3.2 mm long, lacerate below, plicate, essentially naked, yellow to yellowish brown below, yellowish brown or reddish brown above. Spores 16-38 µm, minutely papillose; anisoporous.
Home