Male flowers: calyx (4)5-lobed, imbricate; petals absent; disk glands (4)5, alternating with the stamens, fleshy, contiguous; stamens (4)5(6), opposite the sepals; filaments free; anthers erect, dorsifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode cylindric, not lobed.
Leaves alternate, often long-petiolate, stipulate, simple, entire or toothed, penninerved; petioles bipulvinate; stipules minute and deciduous (or foliaceous and persistent elsewhere).
Inflorescences axillary (or cauliflorous elsewhere), solitary (or fasciculate elsewhere), racemose (or subspicate elsewhere); bracts usually 1-flowered; flowers shortly pedicellate.
Female flowers: calyx as in the male; petals absent; disk hypogynous, cupular, entire; ovary (1)2(4)-locular, with 2 ovules per locule; styles short; stigmas bifid, recurved.
Fruit subglobose or ellipsoid, subdrupaceous, tardily loculicidally dehiscent; pericarp thin; endocarp 1-locular by suppression.
Seeds solitary by abortion, ellipsoid, ecarunculate; testa thin; albumen copious; cotyledons broad, flat, green.
Dioecious trees or shrubs with a simple indumentum.