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Erect or prostrate annual or perennial herbs with 4-angled stems. Stipules connate with the petioles to form a fimbriated sheath. Leaves opposite; blades linear-lanceolate to ovate or broadly elliptic. Flowers not heterostylous or only slightly so, in dense spherical sessile terminal or axillary heads. Calyx-tube obconic, obovoid or subglobose; teeth 4–5, 2 often longer, sometimes with minute supplementary ones between. Corolla salver-shaped or funnel-shaped, the tube often with an internal ring of hairs; lobes 4; throat glabrous or hairy. Stamens inserted in the throat, anthers included or exserted. Disc fleshy. Style short or long, divided into 2 short linear branches. Ovary 2(–3)-locular; ovules solitary in each locule, attached by the middle to the septum. Fruit a thin circumscissile capsule, the upper part splitting off together with the calyx-lobes, the septum persistent. Seeds oblong or globose, the ventral face divided into 4 distinct areas; endosperm fleshy.
Herbs, annual or perennial. Leaves usually narrow, often subsessile; petioles united with the stipules to form a setiferous sheath. Inflorescences of terminal or axillary heads. Flowers minute, 4-merous, the calycine cup short, the lobes unequal; corolla funnel shaped, the lobes 4(-3), valvate; stamens included or somewhat exserted, the anthers dorsifixed. Fruits capsular, 2-celled, each cell 1-seeded, transversely or obliquely circumscissile, the calyx persistent; seeds ellipsoid or ovoid, ventrally 4-lobed and with a cruciform groove.
Corolla salver-shaped or funnel-shaped, the tube often with an internal ring of hairs; lobes 4; throat glabrous or hairy.
Fruit a thin circumscissile capsule, the upper part splitting off together with the calyx lobes, the septum persistent.
Calyx tube obconic, obovoid or subglobose; teeth 4–5,2 often longer, sometimes with minute supplementary ones between.
Flowers not heterostylous or only slightly so, in dense spherical sessile terminal or axillary heads.
Seeds oblong or globose, the ventral face divided into 4 distinct areas; endosperm fleshy.
Ovary 2(3)-locular; ovules solitary in each locule, attached by the middle to the septum.
Leaves opposite; lamina linear-lanceolate to ovate or broadly elliptic.
Erect or prostrate annual or perennial herbs with 4-angled stems.
Stipules connate with the petioles to form a fimbriated sheath.
Stamens inserted in the throat; anthers included or exserted.
Style short or long, divided into 2 short linear branches.
Disk fleshy.
