Male flowers in terminal and subterminal corymbose inflorescences; calyx-tube expanded, usually 4-lobed; petals usually 4, imbricate; disk fleshy, lining the calyx-tube; stamens usually 4, alternipetalous, inserted below the margin of the disk; anthers extrorse.
Capsule ellipsoid, 2-valved, loculicidal; seeds compressed, winged; cotyledons broadly oblong, apex obtuse.
Leaves alternate, simple, usually petiolate.
Dioecious shrub.