Multidentia Gilli

Genus

Angiosperms > Gentianales > Rubiaceae

Characteristics

Shrubs, small trees to 12 m. tall or sometimes pyrophytic subshrubs, mostly glabrous, unarmed; stems often lenticellate. Leaves mostly restricted to new growth, paired or rarely ternate, petiolate; blades chartaceous to coriaceous, typically glaucous beneath, with a conspicuous network of finely, or less often coarsely, reticulate tertiary nerves; domatia present as tufts of hair or absent; stipules sheathing at base, pubescent within, provided with a linear, somewhat keeled lobe. Flowers (4–)5(–6)-merous, usually medium-sized, borne in pedunculate cymes, usually at nodes from which the leaves have fallen, except in pyrophytic species; bracteoles linear to lanceolate, small. Calyx chartaceous or coriaceous; limb cupular, truncate, repand, dentate or cupular below and lobed above. Corolla normal or coriaceous and drying wrinkled; tube cylindrical, subequal to lobes or occasionally much longer, glabrous outside, with a ring of deflexed hairs inside and usually rather sparsely pubescent at throat; lobes reflexed, rounded or obtuse, thickened towards apex. Stamens set at throat; anthers partly or fully exserted, oblong-ovate or oblong, with dark coloured connective tissue on dorsal face except for margin, apiculate or not. Ovary 2-locular, each locule with one pendulous ovule; style slightly longer than or less often up to twice as long as corolla-tube, slender; stigmatic knob ± as broad as long or less often elongate, ribbed, hollow to mid-point, apex cleft ± to mid-point when mature. Disk glabrous. Fruit a 2-seeded drupe, large, subglobose, laterally compressed, often somewhat didymous, often lenticellate or rarely small and didymous (in the West African M. pobeguinii (Hutch. & Dalz.) Bridson), crowned by persistent calyx-limb; pyrenes thickly woody, broadly ellipsoid, sometimes curved, truncate at point of attachment, with line of dehiscence extending from point of attachment to apex then arching on either side back towards point of attachment but stopping short of it, very strongly rugulose (except in M. pobeguinii) see fig. 132, p. 754. Seeds with endosperm entire; testa finely reticulate; embryo slightly curved; radicle erect; cotyledons about 1/3 the length of the embryo, set perpendicular to the ventral face of the seed.
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Pyrenes thickly woody, broadly ellipsoid, sometimes curved, truncate at point of attachment, with a line of dehiscence extending from the point of attachment to the apex then arching on either side back towards point of attachment but stopping short of it, very strongly rugulose, (except in Multidentia pobeguinii); seeds with endosperm entire; testa finely reticulate; embryo slightly curved; radicle erect; cotyledons about one third the length of the embryo, set perpendicular to the ventral face of the seed.
Leaves mostly restricted to new growth, paired or rarely ternate, petiolate; blades chartaceous to coriaceous, typically glaucous beneath, with a conspicuous network of finely, or less often, coarsely reticulate tertiary nerves; domatia present as tufts of hair, or absent; stipules sheathing at the base, pubescent within, provided with a linear somewhat keeled lobe.
Corolla normal or coriaceous and drying wrinkled; tube cylindrical, subequal to the lobes or occasionally much longer, glabrous outside with a ring of deflexed hairs inside and usually rather sparsely pubescent at the throat; lobes reflexed, rounded or obtuse, thickened towards the apex.
Fruit a 2-seeded drupe, large, subglobose, laterally compressed, often somewhat didymous, often lenticellate or rarely small and didymous (in the WestAfrican Multidentia pobeguinii (Hutch. & Dalz.) Bridson), crowned by a persistent calyx limb.
Style slightly longer, or less often up to twice as long as the corolla tube, slender; pollen presenter ± as broad as long or less, often elongate, ribbed, hollow to mid-point, apex cleft to about the mid-point when mature; disk glabrous.
Flowers (4)5(6)-merous, usually medium-sized, borne in pedunculate cymes, usually at nodes from which the leaves have fallen, except in pyrophytic species; bracteoles linear to lanceolate, small.
Stamens set at throat; anthers partly or fully exserted, oblong-ovate or oblong with dark coloured connective tissue on dorsal face except for the margin, often apiculate.
Shrubs or small trees to 12 m tall, or sometimes pyrophytic subshrubs (suffrutices), mostly glabrous, unarmed; stems often lenticellate.
Calyx chartaceous or coriaceous; limb cupular, truncate, repand, dentate, or cupular below and lobed above.
Ovary 2-locular.
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Sexuality hermaphrodite
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Mature height (meter) 12.0
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Hardiness (USDA) 9-12

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