Female flowers: gynoecium of (2)3 or 4 carpels, (2)3-or 4-lobed and (2)3-or 4-locular; carpels ± free above, tapering into ± recurved styles; stigmas ± spathulate, oblong, papillose, lateral, facing inwards; ovules numerous, anatropous, axillary, 2-seriate; no staminodes present
Male flowers: stamens 4 with filaments almost free or (3)4–5(?8) with filaments connate at the base into a central column, anthers basifixed, dehiscing by longitudinal slits, their connective prolonged into a beak; pollen in tetrads; no rudimentary ovary present
Leaves decussate; leaf-lamina digitately nerved, plicate, articulate with the petiole; stipules paired, connate with the petiole, persistent
Flowers sessile in terminal, erect, catkin-like inflorescences; perianth absent; bracts present; bracteoles present or absent
Fruit a capsule, dehiscing by the adaxial sutures of the free apices of the carpels
Seeds numerous, small, pendulous; endosperm fleshy, copious; embryo minute
Dioecious, wholly glabrous, balsamic-resinous shrubs; branches opposite