Male flowers shortly pedicellate; calyx closed in bud, ovoid, apiculate, later splitting into 4–5 valvate lobes; petals absent; stamens many, inserted on a globose receptacle, filaments free, anthers with slightly oblique pendulous thecae, apicifixed, introrse, longitudinally dehiscent, connective produced; disk glands free; pistillode absent.
Female flowers subsessile or very shortly pedicellate; sepals (4)5(6), imbricate, persistent; petals absent; disk hypogynous, annular or cupuliform, entire or more or less crenulate, thick, pubescent or setose; ovary 3-celled, with 1 ovule per cell; styles 3, ± free or connate at the base, bifid or bipartite, papillose.
Leaves alternate, petiolate, stipulate, simple, crenate-serrate to subentire, chartaceous or subcoriaceous, penninerved, sparingly gland-dotted on the lower surface; petioles often pulvinate and geniculate.
Inflorescences axillary, solitary or congested, pedunculate or sessile, spicate, racemose or subpaniculate, unisexual or bisexual; bracts rigid, more or less scarious.
Fruit tricoccous, or dicoccous by abortion, dehiscing septicidally into 3 bivalved cocci leaving a columella; endocarp thinly woody.
Dioecious or monoecious trees with a simple indumentum.
Buds often perulate (furnished with protective scales).
Seeds subglobose, ecarunculate.