Tree, up to 35 m high, dbh up to 60 cm, with up to 2 m high buttresses. Twigs 1.5-5 mm thick, pu-berulous, mostly early glabrescent. Leaves 1-foli-olate to 5-jugate; petiole 1.5-7 cm long, 0.75-1.5 mm thick, terete to semi terete; axes sparsely pu-berulous and mostly glabrescent, or glabrous from the beginning; petiolules 3-10 mm long, above mostly narrowly and deeply, sometimes only slightly grooved, without a median rib but mostly with swollen lateral ribs. Leaflets (narrowly) ovate to elliptic, 4-15 by 1.5-5 cm, index 2.25-4.75, thin-pergamentaceous to coriaceous, glabrous or sometimes beneath along the midrib with a few minute appressed hairs; domatia absent; base acute to rounded, attenuate; sides mostly slightly, sometimes strongly curved, sometimes nearly parallel; apex obtuse or rounded or tapering acuminate, the acumen short, broad, and rounded to long, slender, and acute; midrib above sunken to, sometimes prominulous, nerves 0.5-1.5 cm apart, above prominulous, ± distinctly looped and joined near the margin, nervation rather irregular because of the often great number of variably developed intercalated veins, veins clearly differentiated from veinlets, reticulation very coarse, prominulous at both sides. Inflorescences axillary, pseudoterminal, or terminal. Sepals variably but mostly rather high up connate, 1-2 mm long. Petals absent. Disc glabrous. Stamens (6-)8. Ovary 2-celled. Fruits ellipsoid, 3.25-3.75 by 2.5 cm, glabrous, ± densely set with strap-shaped to filiform, curved, up to c. 15 mm long, appendages, bulbous to triangular at the base, confluent or not; wall hard coriaceous, up to c. 3 mm thick.