Herb to 90 cm, variably pubescent with glandular and non-glandular hairs. Leaves mostly or all radical (cauline leaves occasionally numerous), elliptic, petiolate, the upper ones lanceolate to linear, almost sessile; lamina up to 12 (occasionally 25) cm long; petiole to 7 (sometimes to 14) cm long, narrowly winged. Inflorescence panicle-like, few-branched. Calyx 5–13 mm long, the intersepalar membranes usually conspicuous. Corolla-tube 13–18 mm long, 1–3 mm wide at top of calyx, usually narrowed at base; limb 5–10 (occasionally to 16) mm diam., the lobes notched. Two of upper 4 anthers slightly above the other 2, the filaments to 1.3 mm long; filament of the 5th stamen 4–8 mm long, inserted in lower half of corolla-tube. Cleistogamous flowers occasionally present. Capsule ovoid or ovoid-ellipsoid, 4–9 mm long. Seeds reniform or acutely angled; testa irregularly honeycombed or wrinkled.
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