Herb to 1.5 m; stems and leaves glabrous or variably pubescent with non-glandular hairs, occasionally white-woolly. Leaves usually radical and cauline, ovate or elliptic, the upper ones lanceolate to linear; lamina up to 16 (occasionally to 30) cm long; petiole to 7 (occasionally to 16) cm long, usually narrowly winged, the base sometimes slightly stem-clasping. Inflorescence panicle-like, usually few-branched, pubescent with glandular hairs. Calyx 6–13 (rarely to 26) mm long; intersepalar membranes usually conspicuous. Corolla-tube 17–55 mm long, 2–6 mm wide at top of calyx, distinctly narrowed at base; limb 14–35 mm diam., the lobes obtuse or notched. Upper 4 anthers at about same level, their filaments to 3 mm long; filament of 5th stamen 11–16 mm long, inserted in lower half of corolla-tube. Capsule ovoid or ovoid-ellipsoid, 7–12 mm long. Seeds usually reniform or acutely angled; testa irregularly honeycombed or wrinkled.