Osteospermum calcicola (J.C.Manning & Goldblatt) J.C.Manning & Goldblatt

Multi-stemmed, rhizomatous perennial, 200-300 mm high; rhizome much-branched, ±5 mm diam., producing clusters of tutted shoots that elongate the following year and flower before dying down; stems suberect, sparsely branched, nodding in fruit, glabrous on first season's growth but densely glandular-pubescent on second season's growth, pale green with glandular heads of trichomes purple. Leaves alternate throughout, congested basally, erect, decreasing in size acropetally, lower leaves broadly obovate to oblanceolate, tapering below into long, petiole-like base, midrib prominent abaxially, petiole 35-45 mm long; blade 30-40 x 20-30 mm, leathery, puberulous with glandular and eglandular trichomes, these longer along margins, margins sparsely and weakly ±10-dentate, subobtuse-mucronulate, upper leaves oblong-lanceolate, 10-20 x (3-)5-7 mm. Capitula het-erogamous, radiate, ±60 mm diam., solitary on shortly leafy, peduncle-like stems. Involucre shallowly cam-panulate, 10-12 mm diam.; involucral bracts ±20, uniseriate, lanceolate. 6-7 x 2-3 mm. acute, densely glandular-pubescent, glandular heads of trichomes purple, with scarious margins 0.5-1.0 mm wide. Receptacle flat, glabrous. Ray florets female-fertile, 20-22; tube sparsely glandular-pubescent, ±0.8 mm long, up to 9 setting fruit; lamina spreading but recoiling in afternoon, narrowly elliptic. 4-veined, ±3x as long as involucre, 20-25 x 4-5 mm, straw-yellow marked with dark maroon adaxially in basal 5-6 mm, flushed reddish abaxially, veins red. Anthers vestigial, free, reduced to 4, subulate staminodes ±0.8 mm long, yellow. Style terete, on short stylopodium, branching ±1 mm above mouth of tube, branches narrowly elliptic-lanceolate, obtuse, ±1.5 mm long, yellow, lateral margins stigmatic. Ovary obovoid, 3-angled, ±2 mm long, shortly glandular-pubescent. Cypselae homomorphic, 3-winged, 10-12 x 7 mm, body narrowly turbinate, ±6 mm long, glandular-pubescent, with apical, trifenestrate airchamber, windows ovate, 3.5 x 2.0 mm, wings translucent, ±2.5 mm wide. Disc florets functionally male, numerous; corolla narrowly funnel-shaped, 4-5 mm long, yellow; tube glandular-hairy, ±3 mm long; lobes suberect, triangular, ±2 mm long, with marginal veins joining at sinuses and continuing down tube. Anthers 2 mm long, dark purple with yellow filaments; anther base tailed, tails equalling filament collar; anther appendage ovate. Ovary compressed-ovoid with lateral ribs, ±1 mm long, glandular-pubescent. Style terete, on short stylopodium, shortly bilobed apically, lobes deltoid, acutely papillate with basal fringe of longer trichomes.