Plants medium-sized, yellowish or black [green or brownish]. Stems densely foliate, irregularly branched to subpinnate, branches short, erect, and densely foliate, or elongate, pendent, and terete-foliate; axillary hair proximal cells 1 or 2, short, light brown, distal cells 2 or 3, short. Stem leaves imbricate, lanceolate-acuminate; base ± auriculate; margins plane or recurved distally, papillose-denticulate; costa slender; alar cells differentiated; medial laminal cells short-linear, in rows diverging from costa, seriate-papillose. Branch leaves imbricate; apex sometimes 1-seriate, filiform. Sexual condition apparently dioicous. [Capsule immersed or exserted, oblong-ovoid; operculum rostrate; exostome and endostome papillose, cilia inconspicuous. Calyptra usually hairy. Spores papillose].
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