Stems moderately to lightly pubescent, glabrescent. Leaves with pubescent petioles 5-11 cm; lamina broadly ovate or deltoid-ovate, base cordate or sagittate-cordate (or subtruncate), apex acuminate, 10—16(—23) by 8—12(—23) cm, margin entire or remotely dentate, both surfaces pubescent, usually more densely beneath. Inflorescences axillary, com-posed of a raceme or narrow panicle of cymes, slender, 7-15 cm long, usually densely pubescent. Male flowers on pedicels 1-2 mm: sepals 2 mm long, glabrous, outer ones ± elliptic, inner ones broadly elliptic; petals oblong, 0.75 mm long, flat; synandrium 1 mm long, anthers with transverse slits. Female flowers: sepals subequal, 2 mm long; petals lanceolate, 0.75 mm long; staminodes oblong, 0.5 mm long; carpels ovoid-ellipsoidal, 1.25 mm long with recurved, lobed stigma. Drupes drying irregularly ridged, 8-10 mm long, on pubescent pedicels c. 2 mm, glabrous; endocarp broadly elliptic in outline with conspicuous apical keel, dorsally bearing subadpressed spinules and fimbriate or toothed diagonal ridges with the margin encircled by a skirt-like toothed ridge, the large ventral cavity bordered by a thin incurved toothed margin.