Trees or shrubs, the young branchlets commonly ferruginous-tomentose with stellate or dendroid trichomes, often glabrescent. Leaves petiolate, usually pubescent on the lower surface, the trichomes stellate or dendroid, often appressed and bizonal, the margins entire, crenulate, or dentate. Inflorescences paniculate, axillary or terminal. Flowers bisexual, usually (4-) 5 (-6)-merous, mostly pink or white, umbellate, corymbose, or subcorymbose-racemose; sepals small, open in the bud, connate basally, commonly tomentulose and papillose; corolla rotate, usually tomentulose outside, papillose-tomentose inside at least apically and along the edges, the petals connate basally, valvate, narrow, acutish; stamens (4-)5(-6), inserted near the base of the corolla tube, the filaments well devel-oped, slender to stout, the anthers sagittate, lanceolate to ovate, acute or mucronate or obtuse apically, dorsally punctate or epunctate, dehiscent by introrse slits or apical pores, usually dorsifixed above the base, erect or ver-satile; ovary ovoid or subglobose, the ovules few to numerous, usually 1-seriate, sometimes partially 2-seriate, or rarely pluriseriate on the placenta, enclosed or exposed apically, the style long and slender, the stigma punctiform. Fruit 1-seeded, the endocarp crustaceous, the embryo cylindric, transverse.